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interactions affect the
structure and size of the community
density and dispersion of the population
realm of social behavior
the full range of actions, interactions and communication among individuals and groups
types of biotic interactions
mutualism (cooperation)
predation (parasitism)
competition
commensalism
amensalism
mutualism (cooperation)
both species benefit
predation (parasitism)
one species benefits at the cost of the other
competition
neither species benefit
commensalism
one species benefits, the other doesn’t change
ammensalism
one species is damaged the other one doesn’t change
examples of mutualism (cooperation)
stomach bacteria
pollinators
examples of predation (parasitism)
hawk/mouse
competition can be
interspecific or intraspecific
competition only happens when
it has to because neither one is benefiting from the relationship
liebig’s law of the minimum
growth or success is limited by the resource that is in shortest supply
intraspecific competition
individuals of the same species compete with each other for limited resources
interspecific resources
individuals of different species compete for the same limited resources
patterns of competition
scramble
contest
scramble competition
all individuals share a limited resource, no one gets enough
everyone is negatively affected
contest competition
individuals directly compete for a resource and the winners get access while the losers get nothing
mechanisms of competition
exploitative
interference
exploitative competition
organisms indirectly compete by using up the same limited resource, without directly interacting with each other
interference competition
organisms directly interact to block, exclude or prevent others from accessing resources
examples of exploitative competition
consumption
preemptive
consumption competition
organisms compete by using up the same limited resource, reducing how much is available for others
preemptive competition
an organism occupies a space or resource first and prevents others from accessing it
often without direct conflict
examples of interference competition
chemical
territorial
encounter
overgrowth
chemical competition
an organism releases chemicals that inhibit, harm or kill competitors, reducing their ability to survive or access resources
territorial competition
organisms compete by claiming and defending a specific area (territory), excluding others from its resources
encounter competition
individuals come across each other (encounter one another) and directly compete for a resource at that moment, with one gaining access and the other losing
overgrowth competition
an organisms physically grows over another organism, covering it and blocking its access to essential resources like light, oxygen or nutrients
possible outcomes of competition
species 1 wins
species 2 wins
a draw (stable equilibrium = coexistence)
unstable equilibrium
lotka-voltera model of competition
dN/dt = rN (K-N)/K
only looks at 1 species
population growth equilibrium for one species including competition with species 2
Dn/dt= r1N1 {(K1-[N1+alpha1,2N2]) / K1}
k1 is always on x-axis and k2 is always on y-axis
this generates graphs with what
4 possible arrangements of isoclines
isoclines
lines where there is no increase or decrease in population size
competitive exclusion principle (gauss-paramecia)
2 species that compete for the exact same limiting resource can’t stably coexist
one will outcompete the other → local extinction
resource partitioning allows for
coexistence of similar species
resource partitioning
similar species use the same limited resource is different ways so they don’t directly compete
competition restricts
niches
niche
organisms’ job description
fundamental niches
full range of environmental conditions and resources an organism could use to survive, grow and reproduce if there were no competition or predators
realized niches
the actual conditions and space where a species lives in nature
niche overlap
occurs when 2 species use the same resources or share similar environmental conditions
likely to compete here
niche compression
when a species is forced to use a smaller range of resources (a narrower niche) than it normally would, usually because of competition
competitive release
when a competing species is removed, a species is able to expand its niche and use more resources than it could before
specialists
narrow niche, use a specific range of resources
generalists
wide niche, use many resources
why do predators often focus on the prey that is the better competitor
because there are more of the better competitiors
lotka-volterra predator-prey model
dNh/dt = rNh - eNh - cNhNp
rNh=prey population
eNh=prey dying from natural death
cNhNp=predators eating prey
Nh=prey
Np=predators
big idea of lotka-volterra predatory-prey model
prey increases when reproduction is high
prey decreases when predators are abundant
predator equation
dNp/dt = fcNhNp - dpNp
fcNhNp = predator growth
dpNp = predators death
big idea of predator equation
predators increase when prey are abundant
predators decrease when prey are scarce
key thresholds for prey
prey grows when: Np < r/c
prey decreases when: Np > r/c
key thresholds for predators
predators grow when: Nh > d / (fc)
predators decrease when: Nh < d / (fc)
break down predator prey cycle in 4 parts
prey increasing, predators increasing
prey decreasing, predators increasing
prey decreasing, predators decreasing
prey increasing, predators decreasing
what population lags in predator prey interactions
predator populations
predator responses to prey
reproductive response
aggregative response
developmental response
functional response
developmental response
if you have lots of food as a predatory, you will develop better and faster than if you have limited food
functional response
when there are lots of a certain kind of food, you may become better at handling it
ways to catch food as a predator
chase it down
sit and wait: ambush
use tools
cooperate with others
pros and cons of cooperating with others
pros: more efficient, protection (vigilance), ore time to eat, can’t handle prey along
cons: have to share, increased conspicuousness, have to obtain more food
how do prey species fight back
compensatory response
defensive responses
compensatory response
when a population increases its growth, survival or reproduction after being reduced, helping it bounce back
types of defensive responses
constitutive vs inducible
constitutive defenses
defenses that are constantly present, regardless of whether a predatory is around
inducible defenses
defenses that are activated only when a trait is detected
prey defenses
mechanical
chemical
social
behavioral: alarm calling
camouflage: crypsis
defenses that advertise danger
aposematic coloration
mimicry: batesian vs mullerian
aposematic coloration
bright, noticeable colors that signal the prey is toxic or harmful
batesian mimicry
model: dangerous
mimic: harmless
mullerian mimicry
both species are harmful
special forms of predation
parasitism
cannibalism
true predators
grazers
parasites
cannibalism
eating members of own species to reduce population density and competition/stress
true predators
kill prey quickly and eat a lot of prey over their lifetime
grazers
eat only part of the organism, usually don’t kill it, feed on many individuals
parasites
take small amounts from host, don’t kill host outright, interact with few hosts in lifetime
parasites usually need host for more than just food, they need
a place to live and transportation to the next host
transmission from one host to another of the same species may require a
vector (carrier)
micro parasites
very small parasites that reproduce inside the host and cause disease
bacteria, viruses, protozoans
macro parasites
larger parasites that live in or on a host and do not multiply inside the host
fleas, ticks, worms, plants
holo parasite
completely dependent on their host for nutrients
hemi parasites
can photosynthesize but steal water and nutrients from a host
ecto parasite
parasites that live on the outside of their host’s body
endo parasite
parasites that live inside the body of their host
parasites will
“partition the resource”
definitive host
the host when a parasite reaches sexual maturity and reproduces sexually
intermediate host
the host when a parasite lives and develops but does not reproduce sexually
parasites can be spread by
direct contact (transmission): spend basically whole life cycle with one host
indirect transmission: parasite passed between hosts through an intermediate step, different parts of life cycle are spent with different hosts
facultative parasite
can live with or without a host
parasitism is optional
obligate parasite
must have a host to survive
can’t live independently
temporary parasite
only visits the host briefly (usually to feed)
permanent parasite
lives on/in the host for most or all of its life
kleptoparasitism
stealing food/resources from another organism instead of hunting or gathering for itself
temporary facultative intrapsecific
can lay more eggs
“spreading around” your kids, safer
temporary obligate interspecific
specialized behaviors of young
central place foraging
animal collects food and returns to a home base
costs: search time, travel tome
handling time
time to catch, eat and process food
risk-sensitive foraging
animals consider risk vs reward
risk of predators, risk of not getting enough food
optimal foraging
animals try to maximize energy gained and minimize time, effort and risk
optimal foraging theory
animals behave in ways that max energy gained and minimize costs
marginal value theorem
how long an animal should stay in one food patch before leaving
food in a patch gets used up over time to the rate of energy gain decreases the longer you stay