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Evolution
All the changes that have transformed life on earth from its earliest beginnings to the seemingly infinite diversity that characterizes it today
What is Evolution
Process in which significant changes in the inheritable traits of a species occur over time
Macro evolution
Origin of taxonomic groups higher than the species level. Mainly studies using fossil records
Formation of Fossils (Sedimentary rock)
Richest source of fossils:
Specimens rich in minerals may remain as fossils (Ie. teeth, bones, and shells)
How Fossils Form
Water carries small rock particles to lakes and seas
Dead organisms are buried by layers of sediment, which forms new rock
The preserved remains may later be discovered and studied ‘
Formation of Fossils (Petrification)
Some organic material can be preserved in between rocks and remain preserved through petrification (mineralization)
The process by which organic material becomes a fossil through the replacement of the original material and the filling of the original pore spaces with minerals
Formation of Fossils (Preserved Specimens)
Found in acid bogs, tar pits, permanently frozen ground, amber, or hardened tree sap
When conditions prevent most decomposition, organisms may be preserved nearly intact
Origins of Diversity
The diversity of species → Resulted due to the formation of different species
Charles Darwin proposed the idea of Natural Selection in 1859
Natural Selection
Environment selects individuals with the characteristics that allow them to survive over other individuals with characteristics that are not suited for the environmental conditions
Selected within the variation of the population → selected organisms produce more offspring
Genetic Diversity
Basis of Natural Selection
Changes in the characteristics of a species are produced by combination of
Random genetic mutations
Selection for a particular characteristic which increase survival and breeding success → survival of the fittest
Charles Darwin
Worked as a naturalist
Collected specimens, made observations
Left England believing that species could not change
Alfred Russell Wallace
Came up with a theory very similar to Darwin’s
Not knowing about Darwin’s ideas, he sent a copy of his paper
Darwin quickly wrote: “The Origin of Species”
Galapagos Islands
Key to the development of Darwin’s theory
2 Points made by Darwin
Species were not specifically created in their present forms but have evolved from ancestral species
Described the mechanism of evolution as natural selection
Aristotle’s Views on Evolution
Aristotle believed that Organisms range from simple to complex in a hierarchy → species are fixed and do not change
Ladder of life
Natural Theology
God created each individual species for a particular purpose in nature
Classification revealed the “order” of God’s plan (fixed species)
Jean Baptiste Lamark
Spontaneous generation of species
Species change over time into new species
Inheritance of acquired characteristics
Lamark’s Proposed Mechanism
Use and Disuse: Organs used extensively become larger/stronger, the ones that aren’t use deteriorate
Inheritance of acquired characteristics: Body builder wills on strong/large muscles to offspring
Lamark vs Darwin
Lamark believed that a species could alter it’s characteristics solely based on use and disuse, then pass this on to offspring
Darwin found out that natural selection was the reason for change in characteristics during evolution. Those with favourable characteristics that helped increase survival, were able to live longer and create offspring.
Evidence for Evolution and Natural Selection
Biogeography
Comparative Anatomy
Fossil Evidence
Comparative Embryology
Biochemistry
Biogeography
Geographic distribution of species
Islands have many species of plants and animals found nowhere else BUT are closely related to species of the nearest mainland
Example: Darwin’s Finches on Galapagos Islands
Micro evolution
A change in gene (allele) frequency within a population over time
Evolution on the small scale
Mechanisms of Microevolution
Genetic drift (Bottleneck Effect, Founder Effect)
Gene Flow
Mutations
Non-Random Mating
Natural Selection
Genetic Drift
Changes in frequencies of alleles in a breeding population → occurs due to chance events
Two situations that lead to populations small enough for genetic drift to occur are Founder Effect and Bottleneck Effect
Bottleneck Effect
Natural disasters, disease may reduce size of pop. drastically → small surviving populations unlikely representative of the original population.
Founder Effect
A few individuals from a population start a new population with a different allele frequency than the original population
Gene Flow
Movement of alleles IN or OUT of the population due to the migration of individuals → can introduce new alleles into a population and increase genetic diversity
Mutations
Change in the DNA sequence introduces NEW alleles into a population
Non - Random Mating
Individuals of a certain phenotype choose similar mates OR one with preferred phenotypes
Natural Selection
Alleles being passed on to the next generation in different proportion than the original population
→ due to differential success in survival and reproduction
Types of Natural Selection
Stabilizing Selection
Directional Selection
Disruptive Selection
Sexual Selection
Reminder: Natural selection can affect the frequency of an inheritable trait in different ways depending on which phenotypes in a varying population are favoured
Stabilizing Selection
Acts against extreme phenotypes and favours the more intermediate variants
“Average” = Best Adapted
Directional Selection
Favours an increase OR decrease in the value of a trait from the current population average
Favours relatively rare individuals with more extreme variations of a trait from the average
Occurs during periods of environmental change or colonization of a new habitat
Disruptive (Diversifying) Selection
Environmental conditions vary favours extreme phenotypes over intermediate phenotypes.
Two or more variations or forms of a trait that differ from the current population average
Sexual Selection
Favours the selection of any trait that influences the mating success of the individual
Sexual Dimorphism
Marked differences between the sexes (the result of sexual selection)
Not always adaptive (ie. bright plumage = increased predation)
Therefore, in many cases there is a compromise between the two selection forces
Four reasons why Evolution does NOT Fashion Perfect Organisms
Existing structures are modified rather than scrap ancestral structures and re-design new models
Adaptations are often compromises (Eg. Agile humans → given up structural reinforcement, therefore more prone to sprains, torn ligaments
Not all evolution is adaptive (Chance events can happen)
Natural selection can only edit variations that exist → new genes do not exist on demand