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Raven Bilogy 13th edition, CH 25 : The Origin and Diversity of Life
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Deep Time, Geologic time is divided into
four eons which are subdivided into eras and periods
Deep Time, Geological evidence suggests a meteor hit the earth 4.6 billion years ago, no rocks exist from
Hadean eon (first 500 to 700 million years of Earth’s history)
Deep Time, Geological evidence suggests a meteor hit the earth 4.6 billion years ago, Hadean Earth was pummeled by asteroids
which could potentially vaporize entire oceans
Deep Time, Geological evidence suggests a meteor hit the earth 4.6 billion years ago, when meteor hit debris formed
the moon and the rocky mantle melted as temperatures exceeded 2000° C
Changes in Earth in geological time, CO levels shifted and affected temperature
early atmosphere had high CO levels, water slowly vaporized from molten rock
Changes in Earth in geological time, Increased weathering converted silicate rock to soil
CO formed carbonic acid which released bicarbonate ions and calcium from rock
Changes in Earth in geological time, Decreases in CO₂
lowered Earth's temperature
Continents moved over geological time, Earth’s crust formed rigid slabs of rock called plates
under continents and oceans
Continents moved over geological time, Two supercontinents formed
Rodinia (all continents) and Gondwana (all current Southern Hemisphere continents), Pangea formed from Gondwana
Life emerged in the Archean eon, Proterozoic ("early life") eon occurred
two billion years into Earth’s history, characterized by formation of Rodinia
Life emerged in the Archean eon, Rodinia broke up before Phanerozoic eon
Cambrian period showed diversification of multicellular organisms
Life emerged in the Archean eon, Birds and mammals have existed for
4% of earth’s existence; humans present for 0.2% of earth’s history
Early organic molecules, How first organic molecules formed is not known
hundreds of thousands of meteorites/comets may have carried organic materials, or they originated on early Earth
Early Earth's atmosphere, Few geochemists agree on exact composition
popular view includes CO₂, N₂, water vapor, H₂, and other sulfur/nitrogen/carbon compounds; was a reducing atmosphere
Early Earth's atmosphere, In 1953 Miller and Urey did an experiment
reproduced early reducing atmosphere over water, simulated lightning, temperature below 100°C
Early Earth's atmosphere, Within a week methane gas converted into other simple carbon compounds
which combined to form more complex molecules; later experiments produced amino acids and adenine
Early Earth's atmosphere, Miller and Urey concluded that
key molecules of earth could have formed on early Earth
Evolution of metabolism, Primitive organisms may have been
autotrophic or heterotrophic
Evolution of metabolism, Landmarks in the evolution of metabolism include
oxygenic photosynthesis, carbon fixation, nitrogen fixation
Evolution of metabolism, RNA may have been first genetic material
lipid bubbles could increase probability of metabolic reactions, leading to cell membranes; single-celled organism the first life-form
Conditions on early Earth, It seems likely that Earth's first organisms emerged and lived at
very high temperatures; around 3.8 BYA ocean temperatures dropped to 49-88°C
Fossil evidence of life, Evidence of life during Archean is difficult to find
two main formations of 3.5-3.8-billion-year-old rocks found in Kaapvaal and Pilbara cratons
Microfossils, Microfossils are fossilized forms of microscopic life
oldest are 3.5 billion years old and resemble present-day prokaryotes; fossils from 3.2 BYA could be cyanobacteria
Stromatolites, Stromatolites are mats of cyanobacterial cells that trap mineral deposits
indirect evidence for ancient life, oldest are 2.7 billion years old, modern forms known
Isotopic data, Living organisms incorporate 12C into their cells before other carbon isotopes
analysis of oldest rocks suggests carbon fixation active as long as 3.8 BYA via Calvin cycle or reductive citric acid cycle
Biomarkers, Look for evidence of ancient organic molecules of biological origin
hydrocarbons from fatty acid tails found in ancient rocks, carbon isotope ratios indicate cyanobacteria at least 2.7 billion years old
Earth's Changing System, Climate and atmosphere affect organisms' survival
dramatic shifts led to mass extinctions; Earth has been cooling since formation, extreme drops caused "Snowball Earth"
Shifts in atmosphere, Geological changes explain many changes in atmosphere
hot wet tropics accelerated weathering, Snowball Earth decreased temperature and slowed weathering, plate tectonics can also affect CO₂ levels
Continental motion affected evolution, Continents sit on submerged plates in motion
shifting plates affect evolution by isolating populations or allowing interbreeding
Continental motion affected evolution, Cenozoic era began 66 mya
Australia/Antarctica and Greenland/North America separated, Atlantic Ocean grew, Cretaceous greenhouse conditions submerged continental areas
Oxygenic photosynthesis produced atmospheric O₂, There was a 200-million-year lag between origins of photosynthesis and substantial O₂ levels
iron oxide in oceans, O₂ interacted with UV to form ozone (O₃)
Did plants contribute to glaciations?, Growing evidence that plants contributed to two glaciations
colonization of land followed by glaciation 488-444 mya; vascular plant diversification concurrent with second glaciation 400-360 mya
Ever-Changing Life on Earth, Life evolved into three monophyletic domains
Eubacteria, Archaea, and Eukaryotes
Ever-Changing Life on Earth, Eukaryotes divided into 5 supergroups
Excavata, SAR (Stramenopila, Alveolata, Rhizaria), Archaeplastida, Amoebozoa, Opisthokonta
Compartmentalization of cells, Enabled the advent of eukaryotes
bacteria/archaea ruled for 1 billion years, have less compartmentalization; eukaryotes developed extensive endomembrane system
Evolution of endomembrane system, Infolding of the cellular membrane
formed nuclear membrane (not in bacteria/archaea), allowing physical separation of transcription and translation for added gene expression control
Evolution of endomembrane system, Golgi apparatus and endoplasmic reticulum
facilitate intracellular transport; not all cellular compartments derived from endomembrane system
Endosymbiosis and the origin of eukaryotes, Mitochondria and chloroplasts entered early eukaryotic cells by endosymbiosis
mitochondria descendants of parasite Rickettsia, chloroplasts derived from cyanobacteria
Unicellular body plan, Tremendously successful
unicellular prokaryotes/eukaryotes constitute about half of biomass on Earth, but a single cell has limits with specialization
Unicellular body plan, Multicellularity allowed organisms
to deal with environment in novel ways through differentiation
Multicellularity, Has arisen independently in different eukaryotic supergroups
requires cells to connect and communicate; gene expression varies to allow specialization
Sexual reproduction increases genetic diversity, Allows greater genetic diversity through
meiosis and crossing over
Sexual reproduction increases genetic diversity, First eukaryotes were probably haploid
diploids arose on separate occasions by fusion of haploid cells
Rapid diversification occurred during the Cambrian, Evolutionary innovations occurred while life was primarily aquatic
established foundations for tremendous diversity
Rapid diversification occurred during the Cambrian, Extremely rapid expansion of life called the Cambrian explosion
occurred 542 to 488 MYA; first multicellular animals appeared 50 million years following
Major innovations allowed for the move onto land, Plants and then animals colonized terrestrial environments after Cambrian radiation
evolution of photosynthesis protected organisms, ozone layer protected from UV
Major innovations allowed for the move onto land, Successful movement from water to land required innovations
to prevent desiccation and to obtain wate