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benefits and problems of land
benefits: more light, more available carbon dioxide, and access to mineral-rich soil
problems: desiccation and lack of support against gravity
plants arose from
green algae
closest living relatives of plants
charophytes
similarities of charophytes and land plants
sperm structure, cellulose-synthesizing protein arrangement, and nuclear, chloroplast, and mitochondrial DNA
key adaptations for life on land
alternation of generations, multicellular dependent embryos, walled spores produced in sporangia, apical meristems, cuticles, and usually stomata
bryophytes
nonvascular plants with life cycles dominated by gametophytes, rhizoids instead of true roots, sperm that usually need a film of water to reach eggs, and small, nutritionally dependent sporophytes
ex: liverworts, mosses, and hornworts
two stage life cycle of all land plants
multicellular haploid gametophyte and multicellular diploid sporophyte
gametophyte
haploid stage of plant life cycle that produces gametes
sporophyte
diploid stage of plant life cycle that produces spores
ploidy
number of complete chromosome sets, rather than the amount of DNA alone
haploid
one set of chromosomes (n)
diploid
two sets of homologous chromosomes, one from each parent (2n)
zygote
diploid cell produced by the union of haploid gametes during fertilization
ecological importance of mosses
retain nitrogen, tolerate desiccation, form peatlands, and influence carbon storage and climate
vascular plants evolution and importance
evolved dominant sporophytes, lignified xylem and phloem, true roots, leaves, and sporophylls
important as this allowed plants to grow taller and form the first forests
five major adaptations of seed plants
reduced gametophytes
heterospory
ovules
pollen
seeds
gymnosperms seeds and major living groups
bear naked seeds, typically on cones
major living groups: cycads, ginkgo, gnetophytes, and conifers
angiosperms
flowering plants with flowers and fruits
angiosperms life cycle
pollination, pollen tube-growth, double fertilization, seed formation, and fruit development
evolution
works by modification of earlier structures, not by sudden invention from nothing.
structure fits _____ repeatedly in plant evolution
function
from cuticle to vascular tissue to seeds and flowers
two major problems reproductive success on land depends on
protecting gametes and embryos from drying out
moving sperm to eggs without requiring standing water
angiosperm evolution
includes basal angiosperms, magnoliids, monocots, and eudicots, and pollinator interactions may have contributed to their diversification
algal ancestors favored by natural selection
ones that could tolerate drying at pond margins (trait that helped launch the move onto land)
bryophyte sperm and embryos
sperm: swim through water films to archegonia
embryos: remain protected and nourished in parental tissue
shoot system function
photosynthesis and gas exchange
root system
absorbs water and minerals, anchor the plants, and store nutrients
taproot system
primary root with lateral roots emerging out of it
fibrous root system
no dominant root - dense mat of many tiny roots
root hairs
thin extensions of epidermal cells near the tips of growing roots
root hairs function
increase absorptive surface area
xylem role
conducts water and minerals upward
phloem role
distributes sugars and other organic products
lignin role
stiffens plant bodies for upright growth
roots role
anchor plants and absorb water and minerals
leaves role
increase photosynthetic surface area
endosperm by double fertilization in angiosperms
one sperm fertilizes the egg and the second fertilizes the central cell