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behavior
the internally coordinated response of an organism to internal or external stimuli; the way an organism changes its activity to interact w/environment; has genetic basis → acted upon by selection
tinbergen’s 4 questions
mechanism of behavior (genes, alleles, hormones)
development/ontogeny of behavior (how does behavior change; critical pts)
function/adaptive value of behavior (behavior helps fitness)
evolution/phylogeny of behavior (how older gens contributed to development of behavior)
genetics of behavior
many behaviors are complex traits, thus dependent on many genes—some can be influenced by one gene
development of behavior
some behaviors only expressed at certain times due to changes in gene expression that are mediated by hormones
critical period
sensitive developmental state when learning must take place (ex. imprinting)
how does selection choose individuals to favor?
it favors those with higher fitness benefits than costs and behavioral plasticity
benefits: more food, mates, offspring survival
costs: energetic, risk, opportunity (time)
ecology
study of the interaction between organisms and their environment (biotic and abiotic)
hierarchy of ecological systems
individual, population, community, ecosystem, landscape
difference between weather and climate?
scale; weather is short-term meteorological conditions
climate
large scale w/conditions that occur over long periods of time. shapes environments in which populations evolve and diversify. tropical, arid, temperate, continental, polar
solar radiation’s role
radiation interacts w/atmosphere, oceans, geography to create diverse environmental conditions
greenhouse effect
results from absorption of solar radiation by the Earth and its atmosphere; warms surface and solar radiation turns into infrared radiation that rises and is emitted back down to surface
solar radiation and temp near equator
sunlight travels thru less atmosphere and is spread over a small area
solar radiation and temp at high latitudes
sunlight experiences more atmospheric absorption and is spread over a large area
circulation cells
driven by unequal solar heating, linking temp to precip patterns since warm air holds more water vapor; warm air at equator rises and dry air created at 30 latitude
ocean gyres
caused by earth’s rotation and unequal heating; redistribute heat and shape regional climates
topography
physical features like mountains and plains
rain shadow
caused by mountain ranges that alter elevation and produce distinct climate zones on each side
upwellings
areas of ocean where cold, nutrient-rich water rises to surface along continental coasts
how does proximity to water affect climate patterns?
large bodies of water absorb and release heat slowly, reducing seasonal temperature extremes
how does vegetation affect climate patterns?
forests have low albedo (reflectiveness) because they absorb solar radiation → greater evapotranspiration → precip
how do human activities affect climate patterns?
deforestation increases local temperatures; urban areas create heat islands where buildings and pavement absorb and retain heat
biogeography
scientific study of distribution and diversity of life on earth; species distribution reflects evolutionary history and isolations
why is species diversity is highest in tropics and declines moving north or south away from it?
high speciation and low extinction rates due to less variability in temp and precip, more solar radiation, constant natural selection rates, higher metabolism
tropics had more time to diversify w/o ice ages
higher environmental energy in tropics (supports diversity w/more resources)
population
group of individuals of the same species in a particular area that have potential to interbreed
population dynamics
changes in spatial structure (distribution and density), and changes in growth and size
metapopulations
groups of populations connected by dispersal; species consist of multiple; interconnected subunits of populations
dispersion patterns
uniform — establishing territories
clumped — safety living in group
random — rare, resources are evenly distributed and individuals don’t mind being next to others
what controls population dynamics?
physical environment, biological interactions (competition), dispersal
intraspecific competition
competition for resources by individuals of same species
interspecific interactions
interactions among individuals of different species (predation, competition, mutualism, etc)
dispersal
movement of individuals affects colonization and gene flow; active (animals move), passive (plants use wind to move seeds), facilitated (reliance on another organism to move seeds)
demography
study of changes in population size over time
what contributes to calculating population growth?
births, deaths, immigration, emigration, and population size (N)
simple eqn for closed population size
change in N= rNo where r=(b-d)
what is r?
intrinsic growth rate; large pos values indicate faster population growth
all populations have the potential for ____
exponential growth —when not limited by resources
density-independent factors
factors independent of population size, like natural disasters
density-dependent factors
effects change w/population size, like
limiting resources (intraspecific competition increases w/population size)
predation (higher densities of prey attract more predators)
pathogens (higher densities of hosts = faster pathogen spread)
carrying capacity
K — maximum population size that can be supported by the environment
logistic growth eqn
change in N = rNo(1-N/K) where is N is large, (1-N/K) stops growth
life history
describes the lifetime pattern of growth, reproduction, and survival; strategies evolve thru natural selection on how individuals allocate limited resources and shapes traits of populations
type I survivorship
in many large mammals; high survival rates at most ages that has a steep drop off at old age
type II survivorship
in many birds, fish, plants; steady decrease in survivorship across age groups; impacted by a biotic factor (predator) that impacts them regardless of age
type III
in most insects and annual plants; rapid loss of survivorship that levels off for middle age groups before survivors die
trade offs
when investments in one component of fitness decreases another component of fitness
r specialists
where natural selection favors high growth rates → exponential growth; early reproduction, many small offspring, type III survivorship, fast population growth, high tolerance for environmental variability
k specialists
where natural selection favors carrying capacity → logistic growth; late reproduction, few large offspring, type I survivorship curve, slow population growth, low tolerance for environmental variability
species interactions
affect individuals and populations, can lead to (co)evolutionary change, and vary in direction and strength
coevolution
2+ species affect each other’s evolution (ex. predator-prey); usually result of strong interactions/relationships
predation
predators evolve strategies to capture and consume prey; prey evolve strategies to minimize likelihood of being captured/consumed
carnivory
prey is usually killed; active pursuit, ambush, subdue
prey use structural, behavioral, chemical, and crypsis defenses
herbivory
prey are rarely killed; may drive evolution of structural and chemical defenses in plants
parasitism
hosts are not typically killed; parasites evolve close associations w/specific host species
microparasites
bacteria, viruses, protists — pathogens that cause disease
macroparasites
worms and insects
competition
occurs when two or more species use the same limiting resource
interference competition
direct; active interference w/other species access to resource
exploitative competition
indirect; both species depress each other through shared resource
competitive exclusion
when one species outcompetes another for limiting resources, leading to the local loss of the inferior competitor
competitive coexistence
occurs when species reduce or avoid direct competition, allowing them to persist in the same area
resource partitioning
occurs when species use different parts of the same resource, reducing competition and enabling coexistence
can lead to coevolution over time
character displacement
facilitates coexistence when competing species evolve differences in shared traits where they occur together
increase their differences to reduce competition for resource
mutualism
positive interaction where two or more species benefit from interacting; can be necessary (obligate) or optional (facultative)
how do positive interactions help populations?
they increase population abundance and promote greater community diversity
community
group of species living together at the same place and time; complex networks of interacting species
what are the subsets of species used to define different types of communities?
taxonomic affinity, guild, functional group
food webs
used to represent the trophic or energetic connections among species in a community
species diversity
measure that combines # of species (richness) and their relative abundances (evenness)
community membership
depends on the regional species pool, dispersal ability, abiotic conditions, and species interactions
keystone species
have strong community-wide effects despite to their small size and abundance; may impact abiotic components, in addition to biotic components
foundation species
strong community-wide effects due to their large size and abundance; often habitats/food for other species
ecosystem engineers
create or modify aspects of the physical environment for themselves and thus other species
trophic cascades
occur when strong feeding interactions at one trophic level trigger indirect effects that ripple through the rest of the community
why do more complex habitats support higher diversity?
there are more unique environments, more shelter, and more ways for species to coexist
what type of disturbance shapes community diversity?
communities w/intermediate levels of disturbance support the highest diversity
what happens in communities with high species diversity?
they tend to be more productive, more stable, more resilient (less mortality)
ecosystem
communities of organisms interacting w/their physical and chemical environment, focusing on how energy and nutrients move within environment
primary production
photosynthesis and chemosynthesis; sun is ultimate source of energy
secondary production
consumers biomass (tissue) accumulation
how do nutrients move thru living and non-living environment?
thru soil, water, atmosphere
process by which nutrients are recycled?
consumers eat producers → die → nutrients back into environment → grows producers
how do we measure primary productivity?
using the rate at which light is captured and converted into chemical bonds
gross primary production (GPP)
rate energy is captured and assimilated by products
net primary production (NPP)
rate producer biomass is gained from assimilating energy; supports all life in an ecosystem via production of organic matter
NPP = GPP- respiration
how is NPP affected by climate?
NPP increases w/temp, and increases then declines w/precipitation
net secondary productivity
rate of consumer biomass accumulation; result of consuming primary producers
what limits how long food webs can be?
the fact that most energy is lost from primary to secondary production
how do ocean primary producers differ from terrestrial primary producers?
ocean producers are more r-selected and terrestrial producers are more k-selected
what types of areas tend to have longer, more complex food webs?
areas w/high primary productivity
how long do nutrients cycle?
indefinitely; elements trapped on earth cycling
biodiversity
diversity of life on earth
how do human activities impact biodiversity?
genetic, species, and ecological diversity are declining as a result of human impacts
ecosystem services
instrumental and intrinsic value of ecosystems to humans