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Evolution
 a unifying concept in biology, from the origin and diversification of life to understanding disease and behavior; change in allele frequencies in a population over time
looks at:
changes in trait distributions
why some individuals survive more than others
how populations adapt (or don’t)
Ecology
takes biology from the level of cells and individuals to explain the complexity of interactions between organisms and their environments.
looks at:
change in abundance (population)
interactions b/w species
Ecosystem
a dynamic community of living organisms (biotic component) interacting with their non-living physical environment (abiotic component)
Uniformitarianism
change occurs through continuous, gradual processes over long periods of time
small processes that accumulate
erosion, sedimentation, slow climate shifts, etc.
long timescales
Catastrophism
change occurs through sudden, dramatic events that rapidly alter systems.
large, intense events with immediate effect
volcanic eruptions, earthquakes, floods, glacial events, etc.
short timescales
Trait
any measurable or observable characteristic of an organism.
body size, coloration, growth rate, timing of reproduction, behavior, etc.
Fecundity
the number of offspring an individual can produce
Speciation
the evolutionary process by which populations evolve to become a distinct species
- can happen when gene flow is reduced/stopped
Biological Species concept
a group of individuals that have the potential to interbreed and produce fertile offspring
limitations: fossils, asexual reproduction, self-fertilization)
Gene flow
movement of genes between populations via migration
- followed by reproduction
- keeps populations genetically similar (when a yellow butterfly migrates towards an orange butterfly space, they then reproduce with orange butterflies, and both species become more similar).
- must be disrupted by reproductive barriers for speciation to occur
Prezygotic reproductive barriers (no mating attempted)
Temporal Isolation - they breed during different times of the year
Habitat Isolation - live in different habitats, so they never meet or breed
Behavioral Isolation - unique behaviors, i.e., mating calls
Prezygotic reproductive barriers (mating attempted)
Mechanical Isolation: morphological features prevent successful mating
Gametic isolation: gametes do not unite
Post-zygotic barriers
Hybrid inviability: hybrid development impaired
Hybrid sterility: hybrid cannot reproduce (mules)
Hybrid breakdown: 1st gen hybrids fertile + 2nd gen feeble/sterile
Allopatric speciation
occurs when populations are geographically separated; gene flow is interrupted by physical barriers
i.e., islands, glacial separation, habitat fragmentation
Sympatric speciation
occurs without geographic separation; due to behavioral differences, ecological specialization, and genetic changes (e.g., polyploidy).
Polyploidy
gametes with extra chromosomes due to an error in cell division
change in # of chromosomes → new species
Natural Selection
a process where individuals with certain traits survive and reproduce more successfully than others.
Fitness
an organism’s relative ability to survive and reproduce
high fitness → more offspring
low fitness → fewer offspring
Genetic Variation
difference in genes among individuals in a population
required for evolution
w/o it → no differences in survival or reproduction
Genetic drift
chance events that cause allele frequencies to fluctuate unpredictably from one generation to the next
- Founder effect: individuals isolated from the main population
- Bottleneck effect: population goes through a period in which its size decreases
important in small populations
can reduce/eliminate alleles (reduce genetic variation)
may cause alleles to become fixed
Gene flow
genetic exchange due to the migration of fertile individuals or gametes between populations
reduces genetic differences between populations
increases genetic diversity within a population
can increase/decrease the fitness of populations
Directional selection
favors individuals at one extreme of a phenotypic distribution
population mean shifts in one direction
one trait value is consistently favored
Stabilizing selection
favors individuals with intermediate phenotypes
extremes selected against; good to be average. (high fitness)
variation decreases
population becomes more average
Disruptive selection
favors individuals with extreme phenotypes
low / high = good; middle/average = bad
population splits into two groups
can lead to divergence
Balancing selection
Frequency-dependent selection: the fitness of a phenotype depends on how common it is in the population
common = bad
maintains genetic diversity in population
Heterozygote advantage: natural selection does not always cause the elimination of “weak” or less fit alleles
carrying the recessive gene for sickle cell can protect you from malaria.
Sexual selection
acts on traits that affect reproductive success
mate choice (intersexual selection): one sex prefers certain traits
competition (intrasexual selection)
traits that increase mating success become more common
they may become exaggerated
Artificial selection
humans intentionally choose which individuals reproduce, favoring certain traits.
traits that were beneficial where organisms were bred may not be ideal for the wild.
Macroevolution
major evolutionary changes over a longer period of time
speciation, extinction, large-scale diversification
occurs through extended microevolution over long time scales
Adaptive radiation
rapid diversification of a single ancestral species into multiple species adapted to different environments
occurs when new habitats are available
often follows environmental change
Gradualism
evolution occurs slowly and continuously over long periods of time
Punctuated equilibrium
evolution occurs in rapid bursts, separated by long periods of little change
Homologous structures
anatomical features in different species that share a common evolutionary ancestor
used to reconstruct evolutionary relationships
they are evidence of shared evolutionary history
Analogous structures
biological features in different species that share a similar function but evolved independently (not from a common ancestor)
Vestigial structures
reduced or non-functional features inherited from ancestors
evidence of past environments and evolutionary history
Exponential growth
when resources are abundant and conditions are ideal
population grows faster as it gets larger
Logistic Growth
growth of a population slows over time
resources become limited
competition increases
Carrying Capacity (K)
the maximum population size an environment can support