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Primary growth
Increase in length of stems and roots, driven by apical meristems.
Secondary growth
Increase in girth due to vascular and cork cambia activity.
Apical meristem
Meristematic tissue at root and shoot tips responsible for elongation.
Lateral meristem
Secondary meristems (vascular and cork cambia) that thicken stems and roots.
Vascular cambium
Produces secondary xylem (wood) inward and secondary phloem outward.
Cork cambium
Produces cork cells for bark protection.
Cell division, enlargement, differentiation
Three stages of plant growth: multiplying, expanding, and specializing cells.
Bud initiation and break
Formation and sprouting of buds, linked to seasonal cycles.
Epicormic shoots
Shoots from dormant buds under bark, activated by stress/pruning.
Phenology
Study of seasonal biological events like budburst, leafing, flowering, senescence.
Growth rings (annual rings)
Layers of xylem formed yearly, indicators of age and climate.
False rings
Incomplete or extra rings caused by stress/weather.
Earlywood / Latewood
Earlywood: large, thin-walled cells formed in spring. Latewood: dense, small cells formed later.
Juvenile wood
Early wood near the pith, often weaker and less dense.
Mature wood
Later-formed wood with more stable properties.
Tree crown
The leafy portion of a tree, often classified as dominant, co-dominant, intermediate, or suppressed.
Crown shape
Overall form of the crown (round, conical, spreading, etc.).
Branch phyllotaxis
Arrangement of branches/leaves: spiral, opposite, or whorled.
Branch order
Hierarchical branching: primary, secondary, tertiary, etc.
Apical dominance
Suppression of lateral buds by hormones (auxin) from the terminal bud.
Apical control
Regulation of overall vertical dominance of the main axis.
Monopodial branching
Persistent growth from a main axis (common in conifers).
Sympodial branching
Main axis terminates, and growth continues via lateral branches (common in angiosperms).
Sylleptic branching
Branches develop directly without a resting bud stage (continuous, same year).
Proleptic branching
Branches arise from buds that rested before elongating (delayed growth).
Tree habit
Growth form: excurrent (single trunk, conical; e.g., pine) or decurrent (spreading; e.g., oak).
Root architecture
Arrangement of roots: taproot, lateral roots, adventitious roots, buttress roots.
Canopy layering
Vertical stratification of foliage layers in forests.
Xylem
Water-conducting tissue of tracheids, vessels, fibers, parenchyma.
Phloem
Sugar-conducting tissue with sieve tubes, companion cells, parenchyma.
Tracheids
Narrow, elongated water-conducting cells (main in conifers).
Vessels
Wider, shorter conducting cells forming tubes (in angiosperms).
Rays
Radial parenchyma transporting substances laterally.
Pith
Central parenchyma core in young stems.
Bark
Protective outer covering: phloem, cork cambium, cork, rhytidome.
Heartwood
Inner, non-living xylem, often darker, resistant to decay.
Sapwood
Outer xylem actively conducting water.
Reaction wood
Specialized wood formed under stress: compression wood (conifers) or tension wood (angiosperms).
Lenticels
Small bark openings for gas exchange.
Resin canals
Tubes in conifers producing resin for defense.
Transpiration stream
Flow of water from roots to leaves, driven by evaporation.
Water potential gradient
Force driving water movement in plants.
Photosynthate transport
Movement of sugars from sources (leaves) to sinks (roots, fruits).
Source–sink relationship
Dynamic allocation of sugars depending on demand.
Hydraulic conductivity
Efficiency of water transport through xylem.
Cavitation
Formation of air bubbles blocking water transport.
Non-structural carbohydrates (NSCs)
Sugars and starch stored for metabolism and stress survival.
Carbon allocation
Partitioning of carbon among growth, storage, reproduction, defense.
Secondary thickening
Stem/root girth increase via cambial activity.
Dendrochronology
Tree-ring study for age and climate history.
Leaf area index (LAI)
Ratio of leaf surface area to ground area.
Self-pruning
Natural shedding of shaded/lower branches.
Crown shyness
Gaps where adjacent crowns avoid overlapping.
Root–shoot ratio
Biomass proportion between below- and aboveground tissues.
Root collar
Transition between root system and stem base.
Tree buttress
Flared basal roots supporting tall trees (common in tropics).
Axillary bud
Bud formed in the leaf axil (angle).
Adventitious roots
Roots arising from stems, branches, or leaves.
Growth flushes
Cyclic bursts of shoot elongation (polycyclic: multiple per year; monocyclic: one per year).
Epicormic buds
Dormant buds beneath bark that sprout after stress/damage.
Lignotuber
Woody swelling at base storing buds and carbohydrates for resprouting.