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Order Fagales
Order in the Rosids
Part of Nitrogen-Fixing Clade
Wind-Pollinated Trees
Unisexual Flowers in Catkins
Inferior & Syncarpic Pistils with 2 to 3 Carpels
Nut: Boney One-Seeded Fruit
Family Fagaceae (Oaks & Allies)
670 Species in 7 Genera
Large Deciduous Trees
North Temperate Distribution
Simple Leaves
Nut enclosed by Subtending Bracts
Includes Beeches (Fagus), Oaks (Quercus), & Chestnuts (Castanea)
Family in Order Fagales
Genus Fagus (Beeches)
Genus in Family Fagaceae
Trees with Smooth & Gray Bark
Distributed in North Temperate, Deciduous, & Mesic Forests
Non-Lobed Leaves with Pinnate Venation
Female Flowers with Two Pistils surrounded by One Set of Bracts
Produces Two Nuts

Genus Quercus (Oaks)
Genus in Family Fagaceae
Half of Family
Large Trees with Vertically Ridged Bark
Hybridizing & Taxonomically Challenging Group
Distributed in Woodland, Mesic, Xeric, & Savannah Environments
Deeply Pinnately Lobed Leaves
Female Flowers with Bracts Below
Bracts form Woody Cupule of Acorn
White, Red, & Bur Varieties

Genus Castanea (Chestnuts)
Genus in Family Fagaceae
Trees with Vertically Ridged Bark
American Species Originally Distributed in Central Eastern States
Non-Lobed Leaves with Serrate Margins & Pinnate Venation
Two Female Flowers per Spiny Involucre
Nuts with Fluffy-Looking Wrapper

Family Juglandaceae (Walnuts, Hickory, & Pecans)
Monoecious Trees
Pinnately Compound & Alternate Leaves
Often Aromatic from Resinous Peltate Glands
Allelopathic Defense
Wind-Pollinated & Unisexual Flowers
Nut surrounded by Involucral Bracts that become Fleshy
Family in Order Fagales

Male Flowers of Juglandaceae
Flowers arranged in Pendulous Catkins
Grow on Older Stems
Small Calyx & Absent Corolla
Each Flower Bracted
Three to Many Stamens
Absent Gynoecium

Female Flowers of Juglandaceae
Flowers in Small Groups
Grow on Year’s New Growth
Small & Persistent Calyx often Fused to Involucral Bracts
Absent Corolla
Inferior & Syncarpic Pistil with 2 Carpels
2 Feathery Stigma
Absent Stamen

Floral Formula of Juglandaceae
CA3-6 CO0 A3-♾ OR CA4 CO0 -G-(2)
Examples of Juglandaceae
Black Walnut (Juglans nigra): Tree with Pinnate Compound Leaves & Prized Dark Lumber
Shagbark Hickory (Carya ovata): Tree with Distinct Flakey Bark & Colorful Bud Scales

Family Betulaceae (Birches & Allies)
110 Species in 6 Genera
North Temperate Distribution
Trees & Shrubs
Both Male & Female Flowers in Separate Catkins
Absent Perianth (Calyx & Corolla)
Small Nut or One-Seeded Samara with 3-Lobed Bracts
Family in Order Fagales

Examples of Betulaceae
Birches (Betula): Trees with Pale Bark & Dark Spots
Other Examples: Hazelnut (Corylus), Hop Hornbeam (Ostrya), & Alders (Alnus)

Pepo Fruit
Fruit of Family Cucurbitaceae
Modified Berry with Leathery Rind
Grow on Vines
Examples: Cucumber, Pumpkins, Squashes, Melons, & Gourds
