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taxonomy
science of classifying animals
3 domains
archaea: single-celled prokaryotes, no peptidoglycan, ether-linked
eubacteria: single-celled prokaryotes, yes peptidoglycan, ester linked
eukarya: contain membrane-bound organelles, multicell or unicell
kingdoms
archaea
eubacteria
protist
fungi
plantae
animalia
prokaryotes
no membrane bound organelles
store genetic info in nucleoid region
either archaea or eubacteria domains
Archaea are more closely related to Eukarya than Eubacteria
Archaea possess unique genetic, biochemical, and evolutionary traits that distinguish them from Eubacteria and link them more closely to Eukarya
endospores
preserve genetic material during harsh conditions
extremophiles
Usually archaea
can survive very harsh conditions
peptidoglycan
make up bacterial cell walls
made of sugars and amino acids
capsule of bacteria
protective coating
virulence factor
gram positive bacteria stain color
purple
gram positive bacteria traits
thick, no outer membrane
no LPS
teichoic acid that connects peptidoglycan to the membrane
exotoxins: toxic proteins
gram negative bacteria stain color
pink
gram negative bacteria traits
thin peptidoglycan layer
periplasmic space filled with periplasmic fluid
Lipopolysaccharides (LPS): endotoxins released when bacteria dies
secretes exotoxins
no teichoic acid
prokaryote characteristics
no membrane bound organelles
smaller
no steroids in the membranes
nucleoid region
origine of replication: 1 in eubacteria, many in archaea,
ribosomes 70S (50S+30S)
only flagella, no cilia
glycolysis, pyruvate oxidation, Krebs cycle in cytosol, electron transport chain in cell membrane
eukaryote traits
membrane-bound organelle
cell wall only in plants and fungi
larger
steroids in the cell membrane
nucleus
no plasmids
many origins of replication
ribosomes 80S (60S and 40S)
glycolysis in cytosol, pyruvate oxidation in mitochondrial matrix, Krebs cycle in mitochondrial matrix, electron transport chain in mitochondrial inner membrane
protists
mostly unicellular eukaryotic organisms
1) fungus-like protists
2) plant-like protists
3) animal-like protists
fungus-like protists
lack a cell wall made of chitin
move via cilia or flagella
saprophytic: feed on decaying organic material via phagocytosis
reproduce via asexual and sporulation
plant-like protist
primary producer
produce oxygen
algae have chloroplast and photosynthesis
others are diatoms and dinoflagellates
animal-like protists
protozoa
Food vacuoles with digestive enzymes
amoeba and paramecium
heterotrophic
move via flagella or cilia
often parasitic
Fungi
kingdom under domain Eukarya
mushrooms, molds, yeasts
heterotrophic
saprophytic: eat decay matter
life cycle is haploid
cell wall is chitin
hyphae: branched filaments come together to form a system of mycelium
reproduction: alternate between asexual and sexual (asexual in favorable conditions via budding and sporulation by mitosis) (sexual in unfavorable conditions switch between haploid and diploid phases)
Lichens
symbiotic autotrophs where fungus is paired with algae or cyanobacteria
fungus: protects and provides water and nutrients
algae/cyanobacteria: photosynthesis to provide fungi with food
animalia
eukaryotic, diploid, multicell, heterotrophic, aerobic
phylum: porifera, cnardia, Platyhelminthes, nematode, rotifera, annelida, mollusca, arthropoda, echinodermata, chordata
coelomates
organisms possessing fluid filled cavity called coelom, mesoderm surrounds coelom completely
annelida, mollusca, arthropoda, echinodermata, chordata → most specialized
acoelomates
do not possess coelom
platyhelminthes: least specialized
pseudocoelomates
partially surrounded coelom
nemotoda and rotifera
schizocelomates
coelom begins and splits in mesoderm
Holoblastic cleavage (even), spiral cleavage, determinate cleavage
protostomes: mouth develops first, before anus (from blastopore)
enterocoelomates
coelom begins as lateral out pockets of the archenteron
radial cleavage, indeterminate cleavage, totipotent
deuterostomes: blastopore becomes anus
phylum porifera (under kingdom animalia)
simple, multicell, porous sponges
earliest animal
body asymmetry
parazoan: no tissue organization and no coelom type
sessile: non-mobile
aquatic
cnidaria (phylum under kingdom animalia)
ex: jellyfish, hydra, corals
radial body symmetry
diploblastic: 2 cell layers, ectoderm and endoderm
aquatic
platyhelminths (phylum under kingdom animalia)
ex: flatworms, tapeworms
non-segmented, parasitic worms
bilateral symmetry
cephalization: “sense of head”
triploblastic: 3 germ layers
acoelomate: lack coelom
protostomes
nematoda (phylum under kingdom animalia)
ex: roundworms
non-segmented, parasitic worms
bilateral symmetry
triploblastic
protostomes
pseudocoelomates
rotifera (phylum under kingdom animalia)
non-segmented, aquatic zooplankton
bilateral symmetry
triploblastic
pseudocoelomates
protostomes
annelida (phylum under kingdom animalia)
ex: earthworms, leeches
first to develop segmentation
closed circulatory system: multiple aortic arches and distinct arteries and veins
bilateral symmetry
triploblastic
coelomates
protostomes
mollusca (phylum under kingdom animalia)
ex: snails, squids, octopus
soft bodies, mostly aquatic
bilateral symmetry
triploblastic
coelomates
protostomes
arthropoda (phylum under kingdom animalia)
organisms with segmented bodies and jointed appendages
most diverse phylum, greatest number of species
class insecta: ants, grasshoppers
class arachnida: spiders, scorpions
class Crustacea: lobsters, crayfish, crab
bilateral symmetry
triploblastic
eucolomates
protostomes
exoskeleton made of chitin
open circulatory system: fluid doesnt stay in, hemolymph is fluid, flows into open body cavity called hemocoel
echinodermata (phylum under kingdom animalia)
ex: starfish, and sea urchin
closest to chordates
radial body symmetry
triploblastic
coelomates
deuterostomes: anus forms first from blastopore
chordata (phylum under kingdom animalia)
segmented bodies
vertebrates and invertebrates
bilateral symmetry
triploblastic
coelomates
deuterostomes
chordates
dorsal hollow nerve cord: develops into CNS
notochord: cartilaginous rod derived from mesoderm
pharyngeal pouch (slits): form pharynx, gills, feeding structures, or head and neck
post-anus tail: dissolved in humans
invertebrates
lack vertebrate
ex: lancelets and tunicates
vertebrates (sub phylum) evolutionary path
early jawless fish
chondrichthyes: cartilaginous, jawed fish
osteichthyes: bony fish
lobe-finned fish: bony fish whose fins first resembled limbs
amphibians: first tetrapods, proper limbs with digits
mammals: most evolved vertebrates
class mamalia
closed circulatory system
4-chambered heart (birds also)
lungs
hair
mammary glands (milk)
diaphragms
paleontology
study of fossils
biogeographic evidence
distribution of species around the world
embryology
study and comparison of the developmental stages of embryos in related organisms
comparative anatomy
compares body parts of animals
homologous structures
may or may not perform same the same function but they share an ansestor
analogous structures
same function but not common ansestor
vestigial structures
sever no purpose, homologous to functional structures in other organisms
biochemical methods
DNA sequence comparisons, common pathways like the Krebs Cycle
Catastrophism
mass extinctions in certain areas leads to local populations that are shaped by the organisms that survivied
George Cuvier
Use and Disuse Theory
Jean-Baptiste Lamarok
inheritance of acquired traits: traits acquired through either the use or disuse and then passed to offspring
incorrect theory
Natural Selection Theory
gradual, non-random process where allele frequencies change as result of environmental interactions
random genetic variations amongst individuals can lead to better fitness
Charles Darwin
survival of the fittest
individuals with higher fitness survive and pass on DNA to future generations compared to less fit individuals
fitness: ability to survive and produce fertile offspring
evolution of populations, not individual
Requirements for Natural Selection
Resource demand exceeds supply
variation of traits within a population
variation in traits must be genetically influenced (heritable)
trait variation must be influential for reproduction or fitness
Stabilizing selection
average is favored
normal bell curve
directional selection
one extreme is favored
disruptive selection
rare trait is favored, average is not favored
sexual selection
non-random mating between males and females
females want quality
males want quantity
artificial selection
humans effect the rate of change
Hardy - Weinberg Equilibrium
allele frequency: p + q = 1
genotype frequency: p2 + 2pq + q2 = 1
Hardy - Weinberg Equilibrium Requirements
large population: minimize genetic drift
random mating
no mutations
no natural selection
no migration
microevolution
gene frequency changes over a few generations
non-random mating
mutations
natural selection
gene flow: movement and mixing alleles between populations via migration and inbreeding
genetic drift
random changes in allele frequencies
has a stronger effect on small populations
bottleneck effect
sudden reduction in population size causing a loss of some alleles
founder effect
a few individuals are separated from a large population with different alleles
sources of genetic variation
non-fatal mutations
sexual reproduction
polyploidy: multiple chromosomes introduce variety
balanced polymorphism
maintains a variety of phenotypes within a population
types:
heterozygous advantage: 2 parents produce offspring with better fitness
minority advantage: rare phenotypes offer high fitness. cycles between high and low frequency
neutral advantage: may be beneficial if environment changes
Macroevolution
large scale over a large amount of time
species are reproductively isolates with little to no gene flow between them via prezygotic and postzygotic isolation mechanisms
prezygotic isolation mechanisms
barriers that prevent fertilization
habitat isolation
temporal isolation: reproduce at different times of year
mechanical isolation
behavioral isolation
gamete isolation
postzygotic isolation mechanisms
reduce organisms success of survival after zygote is formed
hybrid mortality: zygote forms but doesn’t survive development or birth
hybrid sterility: hybrid is infertile
hybrid F2 breakdown: F1 generation survives and is fertile, F2 generation does not survive
speciation
how new species are formed
begins with reproductive isolation, populations gradually diverge into two species
allopatric speciation
geographical barriers
adaptive radiation: many species arise from one ancestor to adapt to different environments
specialize to fill different niches within an environment
sympatric speciation
without geographical barrier
due to:
balanced polymorphism: occurs when different phenotypes are maintained within the same area
polyploidy: chromosome duplication
hybridization: hybrids more fit
Phyletic Gradualism
evolution works by small changes over time
FALSE
Punctuated equilibrium
evolution works through short spurts of change with long periods of stasis
divergent evolution
species come from a common ansestor
convergent evolution
unrelated species adapt to similar environments, analogous structures
parallel evolution
species come from a common ancestor and undergo similar changes
coevolution
two species impart selective pressure on each other
hummingbirds and flowers
camouflage
aposematic colorations: warning colors
mimicry
Resembling other species
Batesian: non-harmful species resemble harmful species
Müllerian: two harmful species resemble each other
root of phylogenetic tree
most common ansestor
node of phylogenetic tree
branch point, group splits from one
polytomy of phylogenetic tree
internal node leads to more than 2 tips
clade (monophyletic group) of phylogenetic tree
cluster with an ancestor and all of its descendants
paraphyletic group of phylogenetic tree
an ancestor and some but not all descendants
cladogenesis of phylogenetic tree
divergence of evolutionary lineages from new clades
anagenesis of phylogenetic tree
gradual evolution of interbreeding population without splitting
parsimony of phylogenetic tree
the simpler the evolutionary explanation, the better