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phloem/inner bark
living tissue that transports the sap from photosynthesis passively
created by the vascular cambium
gradient moves from high to low sugar concentration
xylem
transports water up the tree

phloem anatomy
phloem cells are alive and have thin walls
active part has specialized connective cells called sieves
in angiosperms, sieve tubes have companion cells that provide support to cell
in gymnosperms, there are no sieve plates, and no companion cells Gymnosperms have tracheid-like sieve cells that overlap.
why is the phloem a thin layer?
the phloem is a thin layer because not much is made and its typically only active for a year
bark
outermost layer of a woody stem, includes all living/nonliving tissues external to the vascular cambium
what is the purpose of bark
Protection from:
• Bacteria/fungi
• Insects
• Animals
• Some barks contain chemicals
that reduce attack:
• Cinnamon
• Tannins
• Juglone
what is outer bark composed of
mainly old inner bark, phloem, cork is added to waterproof protection

how does outer bark develop
In a young stem, the epidermis acts as the
protective, outer coating.
• The epidermis is later replaced by a corky
layer called the periderm.
• A new meristem (cork cambium) arises on
the outer layer of the phloem.
• Dead phloem cells get “pushed” outward and
incorporated into the bark
how does bark respond to growth
Pholem starts growing before xylem
• Xylem growth puts strain on the phloem
(somewhat absorbed by the expansion of ray
cells)
• Phloem dies, becomes “suberized” by the
cork cambium, and then becomes the
innermost layer of the dead portion of bark
• Outermost layers of bark split or shed.
gas exchange/lenticels
Cork makes the bark mostly
water- and gas-proof.
• But the living cells of the
cambium need oxygen!
• Solution: Lenticels!
why does bark vary so much?
Differences in cork
cambiums
• Periderm characteristics
• Continuous periderm =
smooth or peeling bark
• Growth allocation?
strobeli
gymnosperm cones
angiosperms
produce fruits (seeds
enclosed in an ovary) that arise from
flowers.
gymnosperm
pollen produced and seeds housed in structures called strobil
meristems responsible for bark growth
vascular cambium, cork cambium.
