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While life is diverse there is…
Unity
5 characteristics of ALL biological life
evolve
composed of one or more cells
reproduce
require information
require energy
LUCA
A single-celled prokaryote that all life shares as a common ancestor
Common ancestor
ancestral organism shared by 2 or more descendent linages
Eukaryotic
single-celled
multi-cellular
Prokaryotic
single-celled
Example of a prokaryote
Bacteria
Examples of organisms processing information (5 characteristics of life)
DNA
Respond to stimuli from environment
Maintain internal conditions
homeostasis
Examples of organisms requiring energy (5 characteristics of life)
respiration
photosynthesis
fermentation
When was Earth formed?
4.5 billion years ago
What life was there on earth at first?
single celled
What did first form of life to do the earth?
created O2 leading to the great oxygenation
What did the Great Oxygenation event do?
earth’s atmosphere changed from CO2 to O2, which allowed for the ozone layer to form
What came after ozone formation?
multicellular life
What is the Cambrian explosion?
when major animal life started to form
When did modern humans appear?
200,000 years ago
How many mass extinctions were there?
5
How do mass extinctions happen?
When extinctions occur significantly above the background rate
Global climate or environmental changes are too drastic and/or too fast for organisms to adapt
Mass extinction is not a total extinction
Some life survives and adapts to the new environment so it shapes evolution of the next batch of life
Typological thinking - Plato
until 1700s Plato thought every species was perfect and unchanging
any variation seen in the population is not important
everything was created individually, independently by God
“Plato is Perfect”
Aristotle
Scale of Nature
same as typological thinking but he added organization in ascending order from simple to complex
placed human at the top but below god/angels
“Aristotle is ascending”
Species
a population of interbreeding individuals in nature able to produce viable and fertile offspring but unable to reproduce with other groups
1800s Lamarck
gets rid of perfect idea and instead adds species are changing over time
BUT still kept ladder/scale (change happening linearly)
1850’s Darwin and Wallace (independently)
evolution by descent with modification
species change over time
instead of linear progression, species are related by common ancestry
species change through natural selection
What does evolution explain?
how life changes to produce the diversity of life we see today and the evolutionary past
even with all this diversity there are unifying characteristics of all life (descent with modification)
4 evidence for evolution
geological evidence
fossil record
transitional forms
can be observed right now
Fossil Record as evidence for evolution
through extinction
concept of extinction didn’t exist until 1800 because of typological thinking
99% of all species to have ever existed are extinct
transitional forms as evidence for evolution
intermediate between ancestral and derived (newer) forms
NOT ancestors but relatives (through common ancestry)
Observations for right now as an evidence for evolution
hundreds of populations have been undergoing evolutionary changes
bacteria resisting antibiotics
can be observed on scale of days, weeks, months, depending on species ability to go through generations
evidence for common ancestry
Biogeography and Law of Succession
Homologies
Biogeography
Where we live and why
typological thinking
similar environments = same best lineage for that environment
real life
can see distinct lineages in similar environments
supports descent with modification
Law of Succession
Always see relationship where living organisms today correspond with fossil records in that environment
Types of homologies
structural
developmental
vestigial
genetic
artificial selection
Structural homology
Similarity in anatomy or morphology from common ancestors
usually adult organisms
Developmental homology
similarity in embryotic of developmental stages due to common ancestry
Vestigial trait
structure with reduced (or lost) function but similar to functioning structure related species
Genetic homology
similarity in DNA, RNA, Amino acid sequence due to inheritance from common ancestor
Artificial selection
humans allow only those with particular traits to reproduce
phylogeny
depicts evolutionary relationships
trees show hypothesized evolutionary relationships
Parts of phylogenetic trees
tip/leaves
branch
node
root
time
hashes
Node in phylogenetic trees
the more recent common ancestor shared by 2 or more linages that split
Root in phylogenetic trees
more recent common ancestor for ALL taxa
Ancestor traits
further back in time and shared by a larger group
derived traits
more recent
monophyletic
clade
grouping with a common ancestor and all its descents
non-monophyletic
grouping that lacks a single common ancestor for the group or is missing descendents
Synapomorphies
a trait found in 2 or more taxa that is inherited from their most recent common ancestor and distinguishes these taxa from others
shared derived trait (homologous)
divergent evolution
Process where taxa sharing a common ancestor evolve different traits over time
trait evolved ONCE in common ancestor
homologies → divergent evolution
Homoplasy/analogy
a trait that is shared by 2 or more taxa BUT NOT due to shared ancestry
Convergent evolution
process where taxa share a trait due to evolving under similar conditions NOT from a common ancestor
trait evolved multiple, independent times
homoplasies→convergent evolution
Darwin’s 4 postulates of Natural selection
individuals within a population are variable (variability)
Variability can be passed on to the offspring (heritability)
more offprsing produced than can survive (struggle for existence)
Survival and reproduction are not random (fitness)
Biological Fitness
the ability of the individual to produce surviving, fertile offspring relative to others
adaptation
heritable trait that increases the fitness of an individual in a particular environment
ecology
how organisms interact with each other and their physical environment
Abiotic factors
non living, and can produce selective pressures
terrestrial biomes
combination of climate and vegetation
climate is the long-term temperature and precipitation condition found in area
vegetation is the dominant growth for physical structure
grass, deciduous forest
Marine biomes
characterized not by climate but by depth, light, water movement
vegetation is kelp, plankton, sea grass
Ecologial levels of organisms
individual
population
community - collection of species
ecosystem - biotic and abiotic
global
Predation
one species kills and eat another (prey)
+/-
Herbivory
One species eats a plant
+/-
Parasitism
one species feeds on host, harms it but not instantly kill
+/-
competition
2 or more species use the same limited resource
-/-
mutualism
each species benefits from species interaction
+/+
Keystone Species
species disproportionally large effect/importance on its ecosystem
provide important resource or activity to ecosystem
pollination, predation, beaver dams being built
loss of it HARMS ecosystem
Biodiversity
richness, evenness, and diversity
increases as you go towards equator
Human effects on biodiversity
habitat loss or fragmentation
overexploitation
invasive species
climate change
endangered species
Resource partitioning
when 2 or more species use different recourses, times, space to lower competition
Productivity
rate of new biomass formation by any biological group
sampling effect
more diverse communities = higher change of at least one highly productive species
Resource use efficiency
more diverse species = able to use total resource pool more completely and efficiently than if only one species present
Facilitation
certain species assist the growth of other species by providing them with benefits
mutualism and keystone species
Ecosystem services
all direct and indirect benefits that humans derive from organisms and ecosystem
provisioning services
provide raw materials
food, fuel, medicines, genetic resources
regulating services
part of earth’s life support system
climate moderation, erosion control, water purification
ABIOTIC
Cultural services
enrich quality of life
aesthetics, recreation, education, preserves
supporting services
enable all other ecosystem services
pollination
pest control
nutrient cycling
BIOTIC
Evolution definition
heritable change in trait frequency (or allele) in a population of individuals of some species over time
4 mechanisms of evolution
natural selection
mutation
gene flow
genetic drift
Natural selection
process by which individuals with favorable heritable traits for a particular environment survive and reproduce more
NON RANDOM
can increase, decrease and maintain genetic variation
Mutation
random genetic change
Effect can be neutral, detrimental, or beneficial
SOURCE OF VARIATION
Genetic Drift
RANDOM
decreased genetic variation
Founder effect and bottleneck effect
Founder effect
occurs when new population is established from a small non-representative subset of the original population
Bottleneck effect
a dramatic reduction in population size caused by a random catastrophe
Gene flow
movement of individuals leaving one population, joining another, and breeding in it
INCREASES AND DECREASES genetic variation
RANDOM
African Origins
oldest H. sapiens fossils in Africa
founder effect as evolutionary mechanism as small group migrated each time
bc genetic variation decreased outside Africa
Why do some places in the Americas have high genetic variation?
Gene flow from slave trade/colonization
Evidence that we share common ancestor from Africa
fossil evidence
genetic homology
why are there unique genetic variations in some places?
bc gene flow from neanderthals and others
Natural selection and mutation in different environments
what explains prevalance of rare diseases in populations
Founder effect and LOW gene flow (like the amish) so small population
Endangered species
population size rapidly reduced
genetic variation decreases bc genetic drift
high susceptibility to diseases bc low variation so less chance for beneficial variant existing