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Ch32,33,34,53
(32) Heterotrophy
Obtain energy and organic molecules by ingesting other organisms
(32) Multicellularity
Many have complex bodies
(32) Animals have no cell walls
They lack rigid cell walls and are usually flexible
(32) Tissues
Cells of most animals are organized into structural and functional units called tissues
(32) Tissues- muscle and nervous tissues
Allow for active movement
(32) Active movement
Most move more rapidly and in more complex ways
(32) Sexual reproduction
Most animals reproduce sexually. Diploid phase dominates
(32) Haploid gametes produced by
Meiosis
(32) Haploid gametes
Eggs are non mobile and sperm is mobile
(32) Fertilization results in
Diploid zygote
(32) Embryonic development
Zygote first undergoes a series of mitotic divisions that produce a ball of cells
(32) Body plans
Particular set of morphological and developmental traits, integrated into a functional whole- the living animal
(32) Taxonomists defined the major branches of the phylogeny by the evolution of the following traits:
Presence of tissues (para/eu,eumetazoa)
Symmetry
Body cavity
Embryonic development
Segmentation
(32) Tissues- zygotes are
Fertilized eggs and totipotent
(32) Totipotent
Can give rise to all other body cells
(32) As embryo develops,
Cells specialize
(32) cell specialization is _________ except in ________
Irreversible
Sponges
(32) Sponges
Parazoa
Simplest animal
(32)Parazoa
Defined tissues and organs are not present
(32) Sponges have the ability to
Disaggregate and aggregate their cells
(32) Eumetazoa
All other animals
Distinct and well defined tissues
Irreversible differentiation for most cell types
(32) Symmetry in sponges
Lack any definite symmetry
(32) All other animals have symmetry
Defined along an imaging axis drawn through the animals body
(32) 2 types of symmetry
Radial and bilateral
(32) Radial symmetric
Organisms are also diploblasic
(32) Bilateral symmetry
Evolved at the same time as animals and became triploblastic
(32) triploblastic
3 germ layers
(32) Bilateral symmetry advantages over radial symmetry
Cephalization
Directional movement
(32) Cephalization
Evolution of a definite brain/head area
(32) Germ layers are also known as
Tissue layers
(32) Most animals have embryos that produce
3 germ layers-triploblastic
(32) Triploblastic body
Outer ectoderm (body coverings and nervous system)
Middle mesoderm (skeleton and muscle)
Inner endoderm (digestive organs and intestines)
(32) All triploblastic animals have
Bilateral symmetry
(32) Cnidarians are
Diploblastic
(32) Cnidarians have an
Endoderm and ectoderm
(32) Sponges lack
Germ layers
(32) Body cavity
Space surrounded by mesoderm tissue that is formed during development
(32) 3 basic kinds of body plants
Acoelomates (compact
Hemocoelomates
Coelomates
(32) Acoelomates
No body cavity
(32) Hemocoelomates aka hemocoel
Body cavity between mesoderm and endoderm
(32) Coelomates aka coelom
Body cavity entirely within the mesoderm
(32) Body cavity functions
Fluid cushions the suspended organs
Fluid acts like skeleton against which muscles can work
Cavity enables internal organs to grow and move independently of the outer body wall
(32) Previous phylogenies were based on
The evolution of the coelom
(32) It is now clear that the internal cavity
Evolved many times and is not a reliable character to infer phylogenetic relationships
(32) Coelom appears to have evolved
Once but some clades have reverted to Acoelomates or pseudocoelomate condition
(32) All deuterostomes have
Coeloms
(32) Protostomes are
Variable
(32) Basic bilaterian pattern of development
Blastula
Blastopore
Archenteron
(32) Blastula
Mitotic cell divisions of eggs form a hollow ball of cells
(32) Mitotic cell divisions
Cleavage
(32) Blastula indents to form
A 2 layer thick ball with blastopore and archenteron
(32) Blastopore
Opening to the outside
(32) Archenteron
Primitive gut
(32) Bilaterians divided into 2 groups
Protostomes
Deuterostomes
(32) Protostomes
Develop the mouth first from or near the blastopore
(32) Protostomes anus (if present) develops either
From blastopore or another region of embryo
(32) Deuterostomes
Develop the anus first from blastospore
Mouth develops later from another region of the embryo

(32) Protostomes and deuterostomes
(32) Cleavage patterns of embryonic cells
Spiral cleavage
Radial cleavage
(32) Protostomes cleavage pattern
Spiral cleavage
New cells form to the right or left of previous cells
(32) Deuterostomes cleavage pattern
Radial cleavage
New cells form on top of previous cells
(32) Developmental fate of cells
Determinate and indeterminate
(32) Protostomes cell development
Determinate
Cell fate is determined early
(32) Deuterostomes cell development
Interdeterminate
Cell fate is not determined until after several division
(32) Evolution of segmentation provides 2 advantages
Allows redundant organ systems in adult such as occurs in the annelids
Allows for more efficient and flexible movement because each segment can move independently
(32) Segmentation appeared several times in
Evolution of animals
Anatomy and embryology have been used previously to
Infer phylogeny
(32) Animal Phylogeny sequence data have accumulated
May suggest differnt phylogenetic relationships
Has resolved some problematic issues with previous phylogeny
(32) Relationships among living animals are
Reflected in their phylogeny
(32) All animals share
Common ancestor
(32) Sponges (porifera) are the
Sister group to all other animals
(32) Eumetazoa (true animals) is a
Clade of animals with tissues
(32) Cnidaria
Branch off evolutionary tree before bilateria
(32) Most animal phyla belong to the clade
Bilateria
(32) Clade Bilaterians
All of which are invertebrates, except Chordata, which includes vertebrates
(32) Invertebrates
Animals that lack a backbone
(32) Vertebrates
Animals with backbone
(32) Bilaterians are divided into 3 clades
Deuterostomia
Ecdysozoa
Lophotrochozoa
(32) Deuterostomia includes
Hemichordates (acorn worms)
echinoderms(sea stars and relatives)
Chordates
(32) Deuterostomia clade includes both
Vertebrates and invertebrates
(32) Ecdysozoans and lophotrochozoans are composed
Of entirely invertebrates
(32) Members of ecdysozoa
Secrete external skeletons and as they grow they shed their exoskeleton
(32) Ecdysis process
Ecdysozoa shedding their exoskeleton as they grow
(32) Lophotrochozoa is
Another clade of bilaterian invertebrates
(32) Lophophore
Lophotrochozoans feeding structure
(32) Trochophore larva
Distinct developmental stage of lophotrochozoa
(32) Origin of multicellularity requires the
Evolution of new ways for cells to adhere and signal to each other
(32) Choanoflagellates and animals have sequence similarities in
Genes involved in adherence and attachment
(33) invertebrates
Animals that lack a back bone
95% of known animal species
(33) Phylum porifera
Sponges
Among the most abundant animals in deep ocean
(33) Sponge characteristics
Sedentary or sessile
Parazoa
Asymmetric
Reproduce sexually and asexually
Filter feeders
(33) Parazoa
Lack true tissues
(33) filter feeders
Water comes in Ostia and exits out osculum
(33) 3 functional layers of sponge body wall
Outer epidermis
Mesohyl
Lining of internal cavity
(33) Sponge outer epidermis
Made up of flattened cells
(33) Sponge mesohyl
Middle layer- gelatinous matrix
Spicules
Spongin
Amoebocytes
(33) Spicules
Needles of calcium carbonate
(33) Spongin
Tough protein fibers
(33) Amoebocytes
Transport