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Dark adaptation
Progressive improvement in light sensitivity in low light conditions
Rod are slower to adapt than cones
Red light maintains rods photopigments by preventing bleaching of rods
Trichromatic theory of colour
By Young & hemholtz
Vision is sensitive to primary colours
3 types of cones (blue, green, red)
Colour blindness
Absence or reduced number of one or more cones
Monochromats - no colour vision
Dichromats - Missing 1 cone
Most common in males bc on X chromosome
Opponent processing theory
Phenomenon that trichromatic theory cant explain
After images (seeing opposite colour when you look away bc opponent process cells fatigued)
Why dichromates can see yellow with red or green cone
Dual process theory of perceiving colour
3 pairs of opponent cells (red-green, blue-yellow, white-black)
After images show complement colours
RGC & LGN have some cells respond to red but not green and vice vera
Visual perception pathway
Retina (RGCs) → visual thalmus (LGN) → V1
Different brain areas process different aspects
Size of fovea in visual cortex
Very big, but small in eye. Like motor cortex mapping
Feature detection
By Husel & Wiesel
In cats (similar to human visual system)
Mapped V1
Use minimal patterns to detect object features
V1 cells
Respond to simple features of environment, simple and complex
Higher complexity cells found in higher visual areas (visual association cortex)
Simple V1 cells
Respond best to lines of particular orientation, reduced activity when line id no their preference
Complex V1 cells
Respond to bars over a range of positions
Some only respond to bars in motion
Detect lines and edges
Binding problem
How does the brain bind diverse info to form whole?
Hypotheses - rapid coordinated activity across multiple cortical association areas
Sound
Mechanical vibration. Pitch responds to frequency
Loudness
Determined by amplitude of sound wave. Measured in decibels
Outer ear
Funnels sound waves into ear drum
Pinna and ear canal
Middle ear
Amplifies sound to inner ear
Ossicle bones: malleus/hammer, incus/anvil and stapes/styrup
Inner ear
Has the cochlea to transduce sound
Cochlea
Bony outer part, fluid filled inner part
Has organ of corti and basilar membrane
Basilar membrane
Moves in response to fluid waves
Organ of corti
Embedded hair cells (transducing sound)
Pitch perception
Place theory: a specific place along basilar membrane & A1 match tone with specific pitch
High frequencies base of cochlea, low frequencies apex
Frequency theory
Neurons fire at the frequency of the pitch
100 hz → 100 nerve impulses
Volley principle
Groups of neurons fire in alternating bursts so that the combined firing rate matches higher frequencies
Localization of sound
Locate with respect to body
Brain stem
Monaural cues (distinguish clarity of sound)
Auditory pathway
Cochlea → auditory nerve → brainstem → thalmus (MGN) → primary auditory cortex (A1)
Deafness
Conductive (malfunctioning of ossicles)
Nerve deafness (damage to auditory nerve)
Noise induced (often with tinnitus)
Aging (loss of hair cells & degeneration of auditory nerve)
Smell & taste
Chemical senses
Odours
Air borne chemicals detected by olfactory receptor cells in nasal passage transmit to olfactory bulb in brain
Pheromones
Odourless chemicals for intraspecies communication
Menstrual sync
Is the thalamus involved in olfaction?
No
Olfaction pathway
Odour molecules → receptors → olfactory bulb → limbic system → cortex
Taste buds
Contained in papillae
Separate taste receptors for each of the 5 tastes
Taste happens through activity in neurons in combination of activity in taste receptor cells
All regions of the tongue can detect all taste
Gustatory pathway
Chemical → taset receptor cell → cranial nerves → brainstem → thalamus → gustatory cortex
Somatosensory system
Skin & body senses
Skin (tactile) - touch, temp, pain
Body - proprioception, vestibular
Specialized nerve endings
Found at the end of sensory neurons
Mechanoreceptors - light touch, deep pressure
Pacinian corpuscle - specialized for pressure
Free nervings
Touch, temp, pain
Plentiful
Distributed across skins surface
Disproportionate representation in cortex (small nerve → big area in cortex)
Referred pain
Damage to internal organs caused pain elsewhere in the body
Pain
Sharp, stabbing, aching, ect
Each stimuli has a threshold for causing pain
Harder to localize than touch
Emotional component (S1 & limbic system)
Gate control model of pain
Control pain by controlling thoughts & emotions in reaction to pain
Pain blocked by consciousness - spinal cord acts as a gate
Distractions & pressure effective
Endorphins
Decrease release of pain sensitive neurons
Proprioception sense
How body is positioned, force we are applying, how much limbs are stretched
Embedded in muscle (stretch) & tendon (force)
Sensory neurons → spinal cord → brainstem → thalamus → M1 and S1
Vestibular sense
Which way is up, how fast you’re moving, if you’re moving
Semicircular canals
Not cortically processed
Respond to acceleration
Proprioceptors
Embedded in muscle (stretch) & tendon (force)
Sensory neurons → spinal cord → brainstem → thalamus → M1 and S1