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Spatial distribution (“patchiness”) of resources & female sociality
dispersed resources vs clumped resources
dispersed resources (eg leaves)
no direct competition, little incentive for aggression (bc highly abundant and low quality), scramble competition
scramble competition
females are weakly bonded and often disperse from natal group
scramble competition examples
gorilla (folivorous, leaves everywhere), colobus monkey (leaf eater), guenon (forest monkey, generalist diet)
clumped resources (e.g fruit)
high incentive for aggression, monopolizable, direct competition, contest competition
clumped resources - high incentive for aggression note
bc patchier (found in fewer places & in smaller clumps) and ephemeral (available only for short windows @ a time)
contest competition can happen…
between and/or within groups
between group competition
high population density, low within group competition, communal resource defense between groups
between group competition example
proboscis monkeys
within group competition
low population density, low between group competition, coalitionary support within groups for preferred access to food
within group competition example
baboons
(blank) can limit within group contest competition
fission fusion dynamics limit within group contest competition (spider monkey, chimp, orangutan)
female territoriality
when food patches (like fruit trees) are dense & defensible, females can benefit from keeping exclusive access by defending a territory
female territoriality measurement
Mitani-Rodman “Index of Defensibility” (D) - territorial if D>1, not territorial if D<1
Mitani-Rodman “Index of Defensibility” (D) formula
D = day range (d)/home range diameter (d’)
Female Socioecology & Food
females as the ecological sex