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Population
Group of organisms of a single species that interact and interbreed in a common place (Particular area)
Have to be able to interbreed
Boundaries usually arbitrary
Population Characteristics:
Abundance
Distribution
Composition
Abundance
# of individuals
Density - # of individuals per unit area
Distribution
Spread or scatter
Pattern — e.g. clumped?
Composition
Sex Ratio
The ratio of males:females
Age Structure
The relative proportions of various age groups in a population
Demography
Intrinsic factors that contribute to a population’s growth or decline
“BIDE” Factors

BIDE Factors
Birth Rate (Natality)
Death Rate (Mortality)
Immigration
Emigration
Birth Rate (Natality)
Proportion of a population newly born in a unit of time.
Death Rate (Mortality)
Proportion of a population dying in a unit of time
Immigration
Movement into a given area
Emigration
Movement out of a given area
“Closed Population”
Population where no immigration or emigration occurs
Population change = _____
Births (B) - Deaths (D) + Immigrants (I) - Emigrants (E)
Birth Rate Terms
Fecundity
Fertility
Recruitment
Fecundity
Actual number of organisms produced
Fertility: The POTENTIAL capability of an organism to produce young
Recruitment
Increment of new individuals added to a wildlife population by natural reproduction, immigration, or stocking
Rates
Finite rate = discrete intervals (birth pulse)
Instantaneous rate = time interval very short (continuous) (birth flow)
Finite rate = einstantaneous rate
𝜆 = er
Instantaneous rate = loge finite rate
r = ln 𝜆
Discrete Difference Equations
Models for populations at a specific point in time
Non-overlapping generations/discrete intervals
Birth-pulse
Finite Rates: 𝜆
Continuous Differential Equation
Models that allow for continuous time change in populations
Overlapping generations
Population growth is a smooth curve
Instantaneous Rates: r
Density-Independent Growth
Multiplication Rate Density Independent
Constant
Exponetnial Growth (“Geometric” Growth)
N
Size of population
Nt
# of individuals at time t
Nt+1
# of individuals at time t+1
1 time interval later (usually years)
Equilibrium
B + I = D + E
No
Starting # of individuals in a population at time 0
Nt+1 = Nt(𝜆*𝜆)
Difference equation
use when working step-by-step with consecutive time periods
Nt = 𝜆tNo OR Nt = Noert
Recursion equation
use when projecting far into the future or working directly with an initial population
predicting a future population size
dN/dt
Change in per capita population over change in time
dN/dt = rN
Differential equation
use when modeling continuous exponential population growth under ideal, unlimited environmental conditions
finding the growth rate/speed of change
(ln2)/r = tdouble
Doubling time equation
use when asked “how long will it take for the population to double?”
Population Growth: r vs 𝜆
Continuous/r:
Population Increase: r > 0
Population Static: r = 0
Population Decline: r < 0
Discrete/𝜆:
Population Increase: 𝜆 > 1
Population Static: 𝜆 = 1
Population Decline: 0 < 𝜆 < 1
Exponential Growth Model Assumptions
No I or E
Constant b and d
No genetic structure
No age or size structure
r = _____ on a graph
slope
Death (Two broad categories)
1) Intrinsic (internal)
2) Extrinsic (external)
Intrinsic Death Examples
Genetic factors, organ failure, senescence
Extrinsic Death Examples
Predation, starvation, weather, disease
Density independent
Population processes NOT affected by population size
Density dependent
Population processes affected by population size
Positive or Negative
Population density is at equilibrium only when _______
per capita birth rate = per capita death rate
Environmental carrying capacity (K)
Capacity of the environment to “carry” a population.
Logistic Growth Model Assumptions
No I or E
No genetic structure
No age or size structure
Constant carrying capacity
Linear density dependence
dN/dt = rN(K-N/K) OR rN(1 - N/K)
Logistic growth equation
used to model population growth when resources are limited
Infraspecific variation in mortality rate (it’s not constant!)
Seasonality: mortality rate is much higher during harsh times of the year
Sex difference: males often more vulnerable to predation, females to starvation or predation while birthing or caring for young
Experience: some individuals will be better at avoiding risks
Age: very young and old animals are often most vulnerable, but this varies greatly between species.
Life Table
A table of population data based on a sample of the population showing the age at which each member died
Mortality Schedule
Fecundity Schedule
x
age (time interval)
fx
number alive at age x
lx
proportion surviving from the start of the life table to the beginning of age x
= fx/fo
dx
proportion of original cohort dying during the age interval x to x+1
= lx - lx + 1
Type I Species
Most individuals die late in life
Type II Species
Uniform rate of decline
Type III Species
Huge decline in young
r-selected species (fast)
Boom and bust
Higher r
Type III survivorship
Low parental care
Example: Rabbits, many insects
K-selected species (slow)
Conservative
Lower r
Type I survivorship
High parental care
Example: Elephants
mx
Average number of offspring born per unit time to an individual female of a particular age/Reproductive outlook at age x
Fecundity related
= bx/fx
bx
Number of female offspring produced per age group
Net Reproductive Rate (R0)
mean number of offspring produced per female over her lifetime
Per capita growth rate per generation
R0 = ∑(lx)(mx)
Sums of lxmx
If survivorship 100%, R0 = ∑(mx)
Generation time (G)
Average length of time between when an individual is born and the birth of it’s offspring
Average time for a population to increase by a factor equal to R0
r ≈ (lnR0)/G
If know R0 and generation time, can estimate r
Dynamic Life Table
“Cohort” “Generation” “Horizontal”
Follow cohort of organisms over time
Time-Specific Life Table
“Static” “Stationary” “Current” “Vertical”
Sample of individuals from each age class
Cross-section of population at a specific time
Things that can be gathered from life tables
Survivorship directly observed
Age at death observed
Age structure directly observed
Welfare factors
Necessary aspects for survival
Components of habitat
Food
Water
Cover
Space
Oxygen
Decimating factors
Those that kill directly
Predation
Hunting
Disease
Accidents
Starvation
Welfare Factor — Food
Nutrition: process animals use to procure and process portions of their external environment for the continued functioning of internal metabolism
Forage: Browse and herbage, which is available and may provide food for grazing animals or be harvested for feeding
Food habits: diet of wildlife
Matter
Materials that occur in the physical environment of Earth and its atmosphere
Plants and animals composed of it
Recycled
Energy
Required for growth and all metabolic processes
Not recycled
2nd Law of Thermodynamics
Transformation of energy not 100% efficient
Two consequences:
Less energy at successive levels (10%)
Length of food chain limitied
Rule of 10
Only ~10% passes to the next level. Therefore, ~90% of energy is LOST at each trophic level.
Maintenance Requirements
Intake at which animal’s weight remains constant and the animal remains healthy.
Vitamins
Organic compounds, which occur in food in minute amounts and cannot be synthesized by animals, and are essential for normal life and functioning.
Coprophagy - consumption of feces
Minerals
Essential inorganic elements
Macroelements and trace elements
Geophagy - eating earth/rocks
Salt licks
Function of digestion
Break up large and complex molecules to sizes that may be absorbed and used for metabolism
Two types of stomachs:
Monogastric stomach (humans)
Digastric stomachs (ruminants, multichambered, etc. cows)
Foregut Fermentation
Ruminants
More efficiently digests cell wall
Microorganisms provide nutrients & help detoxify tissues
Processing time slower (70-100 hrs); eat less overall but get more out it
Hindgut Fermentation
Nonruminants
Less efficient type of fermentation
Rapid processing time (30-50 hrs)
Cover
Vegetative and topographic features that provide shelter from weather or concealment from or for pedators
Succession
The orderly progression through time of changes in community composition
Pioneer
The first species or community in succession
Climax
The oldest stage given no disturbance. “ultimate expression” of vegetational development under prevailing local or regional conditions.
Home Range
Area included in the daily, seasonal, and annual travels of an individual
Territory
Part of home range that is actively defended
How is Water Obtained?
Free water
Preformed — acquire the water via the diet (vegetation or prey)
Metabolic — Water is a product of respiration (converting sugar to carbon dioxide and water)
Winterkill
Lakes freeze over, snow falls on top
Eliminates light to plants
Plants and animals exhaust oxygen.
Likely when:
Water shallow
Prolonged periods of ice cover, snow
Decaying organic material (algae, aquatic plants, etc)
Summerkill
Upper layer — photosynthetically active
Lower layer — decomposition, less oxygen
Summer — algae grows
Water temp increases, holds less oxygen
Likely when:
Algal bloom, decaying
Evenings, cloudy days
Lake is stratified and abrupt mixing occurs