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Lineage
A series of ancestors and descendants connected through evolutionary history.
Trait (character)
A feature or characteristic of an organism that can be studied and compared.
Homology
A similarity between organisms caused by inheritance from a common ancestor.
Convergent evolution
The independent evolution of similar traits in different evolutionary lineages.
Descent with modification
The idea that species descend from ancestors but accumulate changes over generations.
Divergence
The process in which related populations or lineages become increasingly different.
Evolution
A change in allele frequencies in a population across generations.
Natural selection
An evolutionary process in which individuals with certain heritable traits have greater reproductive success.
Mutation
A change in DNA that can create a new allele and introduce genetic variation.
Genetic drift
Random changes in allele frequencies caused by chance sampling in a population.
Gene flow
The movement of alleles between populations through migration and reproduction.
Artificial selection
Human-directed breeding in which organisms with desired traits are chosen to reproduce.
Dispersal
The movement of organisms into a new geographic area where they may establish a population.
Biogeography
The study of the geographic distribution of organisms across Earth.
Gradual geological change
The idea that Earth's geological features are shaped by slow processes over long periods.
Principle of succession
The pattern in which fossil organisms in a region resemble living organisms from that same region.
Adaptive radiation
The diversification of one ancestral lineage into multiple species adapted to different ecological niches.
Local adaptation
The evolution of traits that improve fitness in a particular local environment.
Population
A group of individuals of the same species living in an area and potentially interbreeding.
Gene pool
The complete collection of alleles present in a population.
Allele
A version of a gene.
Allele frequency
The proportion of all copies of a gene in a population that are a particular allele.
Genotype
The combination of alleles an individual has for a particular gene.
Genotype frequency
The proportion of individuals in a population that have a particular genotype.
Genotype count
The number of individuals in a population with a particular genotype.
Homozygous
Having two identical alleles for a particular gene, such as AA or aa.
Heterozygous
Having two different alleles for a particular gene, such as Aa.
Dominant allele
An allele whose effect is expressed in a heterozygote under complete dominance.
Recessive allele
An allele whose effect is masked in a heterozygote under complete dominance.
Hardy-Weinberg Equilibrium (HWE)
A model describing genotype frequencies in a population when specified evolutionary conditions are met.
p
The frequency of one allele in a two-allele system.
q
The frequency of the other allele in a two-allele system.
Allele frequency equation
The equation p + q = 1, representing the total frequency of two alleles.
Hardy-Weinberg equation
The equation p² + 2pq + q² = 1, predicting genotype frequencies under HWE.
p²
The expected frequency of the homozygous genotype AA under HWE.
2pq
The expected frequency of the heterozygous genotype Aa under HWE.
q²
The expected frequency of the homozygous genotype aa under HWE.
Expected genotype count
The expected genotype frequency multiplied by the total number of individuals.
Observed genotype count
The actual number of individuals found with each genotype in a population sample.
Chi-squared goodness-of-fit test
A statistical test comparing observed counts with expected counts.
Evolutionary processes
Processes that can change allele frequencies, including mutation, gene flow, genetic drift, and natural selection.
Nonrandom mating
A mating pattern in which individuals do not pair by chance with respect to particular traits or genotypes.
Assortative mating
Nonrandom mating in which individuals with similar traits are more likely to mate.
Disassortative mating
Nonrandom mating in which individuals with different traits are more likely to mate.
Bottleneck effect
A type of genetic drift caused by a sharp reduction in population size.
Founder effect
A type of genetic drift that occurs when a small group establishes a new population.
Genetic diversity
The variety of alleles or genetic differences present in a population.
Fixation
The condition in which an allele reaches a frequency of 1, meaning it is the only allele at that gene in the population.
Loss of an allele
When an allele reaches a frequency of 0 and is no longer present in a population.
Fitness
An organism's relative contribution of viable offspring to future generations.
Heritability
The extent to which differences in a trait are passed from parents to offspring through genetic inheritance.
Phenotype
An organism's observable characteristics, influenced by its genes and environment.
Phenotypic variation
Differences in observable traits among individuals in a population.
Histogram
A graph showing the frequency or distribution of values for a trait.
Directional selection
Natural selection that favors one extreme phenotype and shifts the population mean toward that extreme.
Stabilizing selection
Natural selection that favors intermediate phenotypes and selects against both extremes.
Disruptive selection
Natural selection that favors both extreme phenotypes over intermediate phenotypes.
Population mean
The average value of a particular trait in a population.
Balancing selection
Natural selection that maintains multiple alleles or phenotypes in a population.
Heterozygote advantage
A form of balancing selection in which heterozygotes have higher fitness than either homozygote.
Negative frequency-dependent selection
A form of selection in which a phenotype has higher fitness when it is less common.
Phylogenetics
The study of evolutionary relationships among organisms.
Phylogenetic tree
A branching diagram representing a hypothesis about evolutionary relationships.
Taxon
A named group of organisms; the plural is taxa.
Branch
A line on a phylogenetic tree representing an evolutionary lineage.
Branchpoint (node)
A point on a phylogenetic tree where one lineage splits into two or more lineages.
Root
The base of a phylogenetic tree representing the common ancestor of the groups shown.
Most recent common ancestor
The closest shared ancestor of two or more taxa on a phylogenetic tree.
Sister taxa
Two taxa that share an immediate common ancestor with each other.
Topology
The branching pattern of a phylogenetic tree, showing relationships among taxa.
Branch length
The length of a branch, which may represent time or evolutionary change depending on the tree.
Parenthetical notation
A system using nested parentheses to represent the branching relationships in a phylogenetic tree.
Monophyletic group
A group containing a common ancestor and all of its descendants.
Non-monophyletic group
A grouping that does not include all descendants of its common ancestor or combines taxa without including their proper shared ancestry.
Character transition
An evolutionary change in a trait along a lineage.
Parsimony
The principle of preferring the phylogenetic tree that requires the fewest evolutionary changes.
Nested hierarchical classification
An organizational system that places smaller groups within larger groups based on shared characteristics.
Species
A group or lineage of organisms classified as distinct from other such groups.
Biological Species Concept
The definition of a species as populations that can interbreed and produce viable, fertile offspring.
Morphological Species Concept
A species concept that identifies species using physical form and structural characteristics.
Ecological Species Concept
A species concept that identifies species based on their ecological niche.
Gene flow within a species
The exchange of alleles among interbreeding populations that helps genetically connect them.
Reproductive isolation
The prevention of successful gene exchange between populations through reproduction.
Prezygotic barrier
A reproductive barrier that prevents mating or fertilization before a zygote forms.
Postzygotic barrier
A reproductive barrier that acts after fertilization and reduces hybrid survival or reproduction.
Habitat isolation
A prezygotic barrier in which populations live in different habitats and rarely encounter each other to mate.
Temporal isolation
A prezygotic barrier in which populations reproduce at different times.
Behavioral isolation
A prezygotic barrier in which differences in courtship or mating behavior prevent reproduction.
Mechanical isolation
A prezygotic barrier in which reproductive structures are incompatible.
Gametic isolation
A prezygotic barrier in which sperm and egg cannot successfully fuse.
Hybrid
An offspring produced by individuals from different species or genetically distinct populations.
Hybrid sterility
A postzygotic barrier in which hybrid offspring survive but cannot reproduce successfully.
Speciation
The evolutionary process through which new species form.
Allopatric speciation
Speciation that occurs when populations become geographically separated and gene flow is reduced or stopped.
Sympatric speciation
Speciation that occurs without geographic separation between populations.
Geographic isolation
The physical separation of populations by a geographic barrier or distance.
Essentialism
The idea that species have fixed, unchanging defining characteristics.
Polyploidy
The condition of having more than two complete sets of chromosomes; it can contribute to speciation, especially in plants.
Reproductive isolation evolution
The gradual accumulation of differences that eventually prevents populations from successfully exchanging genes.