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phylogeny
the history of descent with branching
node
in phylogenetic trees, the point where a branch splits, representing the common ancestor from which the descendant species diverged
nodes can be rotated without changing ________________________
evolutionary relationships
phylogenetic trees
provide hypotheses of evolutionary relationships
phylogenetics
the study of evolutionary relationships among organisms
taxonomy
the classification of organisms
phylogenetic tree
a diagrammed hypothesis about the evolutionary history, or phylogeny, of a species
a phylogenetic tree is a ______________________________ of the evolutionary relationships among organisms
reasoned hypothesis
the search for sister groups lies at the heart of _______________
phylogenetics
sister groups
groups that are more closely related to each other than either of them is to any other group
monophyletic group
includes of a common ancestor and all of its descendants
taxon
(plural, taxa) all the species in a taxonomic entity such as a family or genus
paraphyletic group
includes a common ancestor and some, but not all, of its descendants
polyphyletic group
does not include the common ancestor
taxonomic classifications are ________________________ and _______________________
information storage, retrieval systems
classification hierarchy
domain, kingdom, phylum, class, order, family, genus, species
three domains of life
Eukarya, Bacteria, Archaea
homology is similarity by _____________________
common descent
characters
the anatomical, physiological, or molecular features of an organism that varies among closely related species
character states
the observed condition of a character, such as presence or absence of lungs or arrangement of petals
homologous
characters that are similar because of descent from a common ancestor
analogous
similarities due to independent adaptation by different species
shared derived characters enable biologists to reconstruct _______________________
evolutionary history
synapomorphies
shared derived characters
cladistics
phylogenetic reconstruction on the basis of synapomorphies
the _________________ is often favored among multiple possible trees
simplest tree
parsimony
choosing the simplest hypothesis to account for a given set of observations
molecular data complement comparative morphology in reconstructing ________________________
phylogenetic history
phylogenetic trees can help solve ________________________
practical problems
_________________ provide unique information
fossils
fossils provide a selective record of _______________
past life
trace fossils
a track or trail, such as a dinosaur track or the feeding trails of snails and trilobites, left by an animal as it moves about or burrows into sediments
molecular fossils
sterols, bacterial lipids, and some pigment molecules, which are relatively resistant to decomposition, that accumulate in sedimentary rocks and document organisms that rarely form conventional fossils
_______________________ indicate the age and environment settling of fossils
geological data
geologic timescale
the series of time divisions that mark Earth's long history
fossils can contain unique combinations of __________________
characters
rare ____________________ have altered the course of evolution
mass extinctions
mass extinctions
a catastrophic drop in recorded diversity, which has occurred five or more times in the past 541 million years
Tiktaalik
"in-betweener"
Tiktaalik showed mixed traits from each of the major groups ____________ and _____________ it existed
before, after (fins, scales, gulls, wrist bones, fingers, true neck, amphibian skull)
_____________________ and ____________________ complement each other
phylogeny, fossils
agreement between phylogenies and fossil record provides strong _________________________________
evidence of evolution
a _________________________ is a reasoned hypothesis of the evolutionary relationships among organisms
phylogenetic tree
a phylogenetic tree is built on the basis of __________________________________________
shared derived characters
the fossil record provides direct evidence of ________________________________
evolutionary history
phylogeny and fossils provide _________________ and ____________________ evidence of evolution
independent, corroborating
___________________________________ is the key to the biological species concept
reproductive isolation
reproductive isolation is caused by barriers to reproduction ____________ or ________________ egg fertilization
before, after
___________________ underlies the diversity of life on earth
speciation
speciation can occur with or without _________________
natural selection
speciation
the process that produces new and distinct forms of life
species plot
a graphical representation of characteristics of individuals
species
the fundamental evolutionary unit
______________ are reproductively isolated from other species
species
biological species concept (BSC)
the concept that species are groups of actually or potentially interbreeding populations that are reproductively isolated from other such groups
the ____ is the most widely used and accepted definition of species
BSC
The BSC is more useful in __________ than in practice
theory
morphospecies concept
the idea that members of the same species usually look like each other more than like other species
the BSC does not apply to ___________ or __________ organisms
asexual, extinct
_____________ and __________ complicate the BSC
ring species, hybridization
ring species
species with populations that are reproductively but not genetically isolated
hybridization
interbreeding between two different varieties or species
limitations of the BSC and mophospecies concept
1. difficult to apply in real world
2. cannot be applied to asexual or extinct organisms
3. does not account for genetic exchange in ring species
4. does not account for hybridization in plants
____________ and _____________ can extend the BSC
ecology, evolution
ecological niche
a complete description of the role a species plays in its environment
ecological species concept (ESC)
the idea that there is a one-to-one correspondence between a species and its niche
phylogenetic species concept (PSC)/evolutionary species concept (EvSC)
the idea that members of a species all share a common ancestry and a common fate
___________________ isolating factors occur before egg fertilization
pre-zygotic
___________________ isolating factors occur after egg fertilization
post-zygotic
pre-zygotic isolating factors
behavioral, physical, temporal (time), ecological (space)
post-zygotic isolating factors
genetic incompatibility, sterile offspring
behaviorally isolated
individuals mate only with other individuals based on specific courtship rituals, songs, or other behaviors
gametic isolation
incompatibilities between the gametes of two different species
mechanical incompatibility
structural configuration of the genitalia that prevents mating with another species
temporal isolation
pre-zygotic isolation between individuals that are reproductively active at different times
geographical isolation
spatial segregation of individuals
ecological isolation
pre-zygotic isolation between individuals that specialize ecologically in different ways
genetic incompatibility
genetic dissimilarity between two organisms, such as different number of chromosomes, that is sufficient to act as a post-zygotic isolating factor
speciation is a by-product of the genetic divergence of ________________________
separated populations
partially reproductively isolated
describes populations that have not yet diverged as separate species but whose genetic differences are extensive enough that the hybrid offspring they produce have reduced fertility or viability compared with offspring produced by crosses between individuals within each population
______________ speciation is speciation that results from the geographical separation of populations
allopatric
allopatric
"different place"
allopatric populations
populations that are geographically separated from each other
subspecies
allopatric populations that have yet to evolve even partial reproductive isolation but which have acquired population-specific traits
_______________ and ________________ can isolate populations from each other
dispersal, vicariance
dispersal
the process in which some individuals colonize a distant place far from the main source populations
peripatric speciation
a specific kind of allopatric speciation in which a few individuals from a mainland population disperse to a new location remote from the original population and evolve separately
adaptive radiation
a period of unusually rapid evolutionary diversification in which natural selection accelerates the rates of both speciation and adaptation in a single lineage
_______________ is speciation that occurs in response to speciation in another species
co-speciation
co-speciation
a process in which two groups of organisms speciate in response to each other and at the same time
sympatric populations - those not geographically separated - may undergo ________________
speciation
sympatric
"same place"
speciation can occur ________________
instantaneously
instantaneous speciation
speciation that occurs in a single generation
polyploidy
multiple chromosome sets
most speciation is thought to happen ________________
allopatrically
variance-derived speciation
a change in the physical environment causes allopatry
peripatric speciation followed by _______________
adaptive radiation
natural selection does not always lead to __________________
speciation
reinforcement of reproductive isolation (reinforcement)
the process by which diverging populations undergo natural selection in favor of enhanced pre-zygotic isolation to prevent the production of inferior hybrid offspring