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Reproductive Isolation
the existence of biological factors (barriers) that impede members of two species from interbreeding and producing viable, fertile offspring
impede us from knowing if they are a species based off of the biological species concept that says a species is a species if it has the potential to interbreed and produce viable offspring
-so therefore it is a roadblock to new species because it hinders interbreeding
-blocks gene flow which hinders the two populations to become more alike each other and doesn’t allow for hybrids to come about
Biological Species Concept
a species is a group of populations whose members have the potential to interbreed in nature and produce viable, fertile offspring—but do not produce viable, fertile offspring with members of other such groups
speciation can occur: by the evolution of reproductive isolation
Ecological Species Concept
defines a species in terms of its ecological niche, the sum of how members of the species interact with the nonliving and living parts of their environment (see
Morphological species concept
distinguishes a species by body shape and other structural features
Prezygotic Barriers
a barrier that hinders the breeding process between two species can or prevents the completion of the fertilization process if mating does occur
Habitat Isolation: when species occupy to different habitats so they don’t even come across each other to be able to breed
Temporal Isolation: when species breed during different times, seasons, years
Behavioral Isolation: when there is a difference in courtship rituals
Mechanical Isolation: when two species are unable to interbreed due to differences in morphological structures
Gametic Isolation: when the sperm of one species is unable to fertilize the egg of another species
Postzygotic barriers
stops a hybrid individual from becoming a viable and fertile adult
Reduced Hybrid Viability: the genes of the parents combine in a way that hinders the hybrid's development and ability to survive in the environment
Reduced Hybrid Fertility: different numbers from the parent species doesn't allow it to to produce offspring
Hybrid Breakdown: that current hybrid is viable and fertile, but when they mate the next generation is not
Allopatric Speciation
Speciation that occurs due to a geographical barrier that separates a population into subpopulations
The divide leads to accumulation of differences caused by mutations, natural selection, and genetic drift
This leads to reproductive isolation
This can be caused by mountain ranges coming up or rivers splitting off into 2 streams
Or migration onto a far-off, secluded island [ a prexisting barrier] where genetic drift and founder effect occur strongly( due to smaller size of the population)
Galapagos Finch: they colonized an island and they are separated geographically and then they diverged into multiple different species; each adapted to a different local environment
Sympatric Speciation
speciation that occurs in populations that live in the same geographic area
can occur if gene flow is reduced by such factors as polyploidy, sexual selection, and habitat differentiation (when a subpopulation exploits a habitat or resource not used by the parent population.).
Plants are mor likely to go through sympatric speciation due to polyploidy (a mutation that causes an organism to have more chromosomes than normal). Plants can go through this and survive while it is not as likely for animals. This is because they can produce asexually
Evolution
the accumulation of differences from their ancestors as they adapt to different environments over time
change in allele frequency
Change in a population’s heritable traits or gene pool over time
Homologous Trait
a trait that is shared between two species because they share a common ancestor
Convergent Trait
A trait that two species share because it evolved multiple times separately
Came about independently
Lineage
ancestors and their descendants over time. all the species between the common ancestor and the organism we are focusing on
trait
a characteristic that can be measured or observed
Descent with Modification
the pattern of evolution
Descent: common ancestor
Modification: there is a change in heritable traits and this occurs independently
Divergence: the accumulation of differences(mutations, genetic drift, different selected traits due to different environmental pressures) and they become different species
As the lineage evolves through time they become more like modern species
They look less like each other and their common ancestor
They diversify into multiple species9doversification)
Struggle for existence
many offspring die before they can reproduce
which is why big growth potential is only potential because of limiting factors(predation, habitat loss, pathogens, pollutions)
Natural Selection
occurs when there is variation in a trait that improves the likelihood of survival and reproduction; a trait where those who have it are more likely to survive and reproduce
does each animals have the same likelihood of surviving no? a trait has variation and if it helps survival then naturally those who do not have it die, and then it gets selected and evolves in the species
changes populations not individuals(it is filter) your not gonna see the effect in one individual because it is an average and it occurs over time
changes allele frequencies, increasing the frewuency of advantageous ones and decreasing the non advantageous ones
Evolution by natural selection occurs when there is variation in a heritable trait
Genetic Drift
random events that drastically change the size of a population: habitat destruction, wildfires, (bottleneck effect) ; changes allele frequencies rather they are advantageous or not
Gene Flow
when a group from a population either moves in or out of an area bringing with them new alleles or removing them when they leave. this changes allele frequencies leading and creates diversity
Non-random mating
when individuals choose their partners based off of specific traits, inbreeding, genotype prevalences change in the population ; changes the percentages of homozygotes and heterozygotes in a population; alters genotype frequencies rather than allele frequencies so it is not as strong as genetic drift or natural selection
Darwin’s argument for natural selection
Observation #1: individuals of the same population have varied inherited traits
Observation #2: individuals produce more offspring then the environmental can sustain so not everyone survives and reproduces
Inference #1: if have an inherited trait that increases your likelihood of survival and reproduction you are more likely to have more survived offspring
Inference #2: This unequal ability for an organism to survive will lead to the accumulation of favorable traits in the population over time
Darwin’s Theory of Evolution
Pattern: Descent with Modification
Process: Natural selection
England and Europe Beliefs 1800s
special creation- God created each individual animal and as they are was how they were made; they didn’t descend from anything
The earth is older than 6000 years old but don’t know exactly how old
Paleontologists found fossils so they knew there was change over time(pattern) but the process that were causing it
Darwin’s observations and finds
Argentina
Darwin found a fossil of an extinct big mammal and so that it was very similar to present day animals that occupied the area
Principle of Succession: animals in a region look similar to fossils of that region because that is who they descended from with modification
Chile, Andes Mountains
fossilized seashells at 12,000ft
Earthquake and volcanic eruption raised the coastal shelf 8ft
Charles Lyell- if given enough time these big structures can be made then maybe over time life can change over time
An old earth means there was enough time for descent with modification to generate the entire tree of life
DIVERGENCE
Galapagos Islands
Islands with sparse tall vegetation had tortoises with bumpy shells and long necks/legs
Islands with thick, low vegetation had tortoises with short necks/legs and soft shells
Islands with intermediate features had intermediate featured tortoises
each had features that were best suited for their environment
Different selection pressures lead to local adaptions by natural selection which lead to more branches on a tree(speciation)
Charles Lyell
Hypothesis for gradual and consistent geological change
The small, observable forces we see today, have caused the big things we see today given enough time. Therefore the earth must be very old becuase we got these big freaking mountains; erosion, sedimentation…
Artificial Selection
the creation of different types of an animal by going from the common ancestor and selecting certain traits
what is desired by humans while natural selection is chosen naturally for an organisms survival and reproduction
How it helped Darwin: If I can breed two things and drastically change their characteristics in a short amount of time, nature can co the same over a long span of time. but for natural selection, nature acts as the filter rather than a human being it for artificial selection. He also noticed that when breeding the offspring naturally inherit from the parent
Adaptive Radiation
when species from a common ancestor diversify(or speciaize) in order to fill different ecological niches. It is a burst of evolution that gives rise to lots of new species: Galapagos Finches and Lemurs
speciation is when one population splits off into 2 reproductively isolated populations
Local adaptation to different environments contributes to divergence between populations and the origin of new species. When this happens repeatedly in one lineage, the result is an adaptive radiation: one species splitting into many species.

Origin of Species Ideas
All organisms evolved from a common ancestor via descent with modification and divergence
The mechanism was natural selection
Local Adaptation
Differential Selection Pressures: different environments favor different traits
Microevolution
smaller scale and we can see it in our lifetime
genetic drift, gene flow, mutation, natural selection
changes in the gene pool
Galapagos beak size, antibiotic resistance
Macroevolution
on a bigger time scale
origin of new species
major changes in traits
evolution, adaptive radiation
Principle of Stratigraphy
sedimentary rocks, where fossils are found, the most recent animal will be found on top while the older more archaic will be found deeper on a lower level
Sedimentary Rock Formation
weathering: baby rocks and stuff
erosion: wind and water move these sediments to nearby lakes and river where they settle down
compaction: other sediments(with animal remains) come and compact each layer cause the weight is heavy and as they do this water and air get pushed out and each layer gets cemented together
plant and animal remains become fossilized over time
A fossil rabbit in the Precambrian
this would challenge the idea of descent with modification because there is nothing in the Precambrian for a rabbit to be descended from. It would break the pattern: species evolving over time into other similar species
we haven’t found this but wehave found transitional fossils
Fossil Record: evidence of evolution
Fossil record with lower strata showing more simpler animals and higher strata showing more complex and more modern organisms Extinction: these animals died and leave behind empty ecological niches giving room for other species to enter and diversify. Animals appearing and disappearing align with environmental pressures showing that macro natural selection is occurring. Shows that life does change over time.
Transitional Forms: these are intermediate figures that connect an ancestor to its descendent. It reveals that there isn’t a gap or jumping going, life is changing continuously in connection. It shows descent with modification is taking place
Biogeographical evidence: Darwin’s 3 places he visited
TikTaalik
transitional form that was found
Tetrapod Similarities: Flat head with eyes on top as well as ribs
Fish Similarities: Scales and gills
Tetrapod
4 limbed vertebrae anima; back bone and limbs with digits; amphibians , mammals, reptiles
example: lizards
Archaeopteryx
beak and wings like bird and clawed forelimbs with boney tail like reptile
Vestigial Structure
like vegetable state; a trait that is functionless or almost functionless in an organism(functions in another species) but is still present; it was functional in the common ancestor
example: appendix(help digest cellulose), tailbone(a tail to help with balance), wisdom teeth(grind down on tough uncooked foods)
They show that we share common ancestry with those organisms that had the trait fully functioning. It shows that we inherited things from their ancestor
example: Whales have small pelvic bones because their ancestors had hindlimbs they used to walk on land. That remaining pelvic bone is proof they evolved form their ancestor
Anatomical and Molecular homologies
structure is similar due to shared ancestry but functions differently due to modification
Humans use forelimbs for grabbing while whales use them for swimming; this shows that we evolved from a common ancestor though are functions are different because the structures are the same
Developmental hoomology
inherited similarities during development phase though very different lookin gin adulthood
embryos of fish and chicken have the same tails; inherited developmental features from a common ancestor
Molecular Homology
all organims all have the same codons coding the same amino acid; similarities on the molecular level
Humans and chimpanzees share many similar DNA sequences which shows that they inherited genetic information from a common ancestor
Evolution in the past 50 years
Pesticide resistance: general bug killer used and many but due to mutations not everybody died. So those who survived passed down their resistance and the population evolved and developed this resistance
Evolution in genetic terms
change in the gene pool over time
Gene
a small region of a chromosome that codes for a molecule
Allele
versions of a gene
Diploid
have two copies of gene
Genotype
the allels and individual has for a gene
Population
a group of interbreeding individuals
Allele Freequency
all copies of a given gene that have that allele
p+q=1
Microevolution
change in allele frequencies over time
Hardy-Weinberg Equilibrium
when a population is not measurably evolving caused by:
No genetic Drift
No mutations-source of all genetic variation but is a very slow process
No gene Flow- alter allele frequencies in a population by bring in new ones; they reduce genetic differences between the two populations but introduce variation to a genetically drifted population; goes against NS by bringing in non adapted or selected alleles
No Natural selection- can select for one genotype against others, changing allele frequencies
Random Mating(no inbreeding)-(non random effects genotype frequencies; can produce excees homozygotes(assorttative mating) or excess heterozygotes(disassortative mating)))
the purpose of the test is to see if any of this is happening
Genotype Count
all the individuals in thepopualtion that have the genotype
Genotype frequency
percentage of individuals that have a certian genotype
Gene pool
all genes and alleles in a population
Genetic Drift
change in allele frequencies due to random sampling effects
it is random what alleles will be passed down to offspring, it is random which individuals survive and reproduce
This a loss of genetic variation because as time goes on because of loss of alleles or alleles may become fixed
It is always happening but it is stronger in small populations
works against natural selection as it is random while natural selection is intentional. trying to select the favorable traits and your deleting animals that may have them. less variability for it to be able to select
occurs in every population of any size
Genetic drift arises from sampling error when a finite population passes genes from one generation to the next.
Genetic drift is simply a "sampling error" in how alleles get passed down
Gene Flow
migration, introudicing new allelle or already presetn not as stron alleles to a population. Makes that population similar to the one it came from. Works agaisnt natural selecction as it brings in no adapted alleles.
example: florida pantether was dying so they reintrouced a couple more to the environment which spruced up genetic diversity, brought in new alleles
Mutation
the source of all genetic variation(creates entirely new alleles) and therefore VERY important for evolution and is the permanent change in a DNA sequence(caused by copying errors or environmental factors like radiation)
Mutation rate is too low to have an effect on allele frequencies in the short term and it also rarely happens because there are checks during the copying process that hinder a mutation from becoming permanent; therefore this is probably not a reason for a population to depart from Hardy-Weinberg Equilibrium
Assortative mating
mating based off of similar traits that the other organism has; this increases homozygotes in a population[nonrandom mating]
Disassortative Mating
mating based off of your differences with the other organism; increases heterozygotes in a population[non-random mating]
phylogeny
the evolutionary history of a species
Taxon
a named group on any level of the hierarchy; Panthera is taxon on the genus level and Mammalia is taxon on the class level
Sister Taxa
a group of organisms that share the same common ancestor that others do not share
Clade
a single common ancestor and all its descendent
Parsimonous
the fewest evolutionary events
Topology
branching pattern of the tree; shows you which organisms are more closely related
Sexual Selection
Based off of offspring sometimes at the expense of survival; selection for greater reproduction
Intrasexual selection: Competition between males for mates
Intersexual selection: Choice of the females for males
Leads to sexual dimorphism where the males and females look different
Half-life
the amount of time it takes for half of the atoms of the parent isotope to decay into the daughter isotope
Radioactive Isostopes
decay into other isotopes or elements at a constant rate
Carbon dating:(also 14C) Carbon14 to 14N; the clock starts at DEATH
half-life of 14C is 5730
each half -life is not a lot of time; carbon decays really fast so for those animals who are more than years old, carbon 14 isotope can’t be used for older, older, fossils
for recent organic
Uranium Dating: Uranium 235 and 238(704 my and 4.5 by)
for old inorganic
Clocks: magma or lava cools and solidifies into rock. the U gets trapped during crystallization, beginning the clock