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Cuticle
Waxy coating reducing water loss.
Epidermis
Protective upper/lower cell layers, often with cuticle.
Trichomes (leaf hairs)
Hair-like outgrowths from the epidermis; reduce water loss, reflect radiation, deter herbivores.
Pubescence
General term for hairiness of leaves; adaptation to heat and water stress.
Stomata
Pores in epidermis allowing gas exchange.
Guard cells
Cells controlling stomatal opening and closing.
Mesophyll
Internal leaf tissue specialized for photosynthesis, includes palisade and spongy layers.
Palisade mesophyll
Columnar cells below upper epidermis, rich in chloroplasts; main site of photosynthesis.
Spongy mesophyll
Loosely arranged cells with air spaces aiding gas exchange.
Veins
Vascular bundles containing xylem and phloem.
Veins (vascular bundles)
Contain xylem and phloem for water, nutrient, and sugar transport.
Bundle sheath cells
Cells surrounding vascular bundles, important in C4 photosynthesis.
Chloroplast
Organelle where photosynthesis occurs.
Grana
Stacks of thylakoids.
Thylakoids
Membrane sacs where light reactions occur.
Stroma
Fluid inside the chloroplast where Calvin cycle occurs.
Bulliform cells
Large epidermal cells that facilitate leaf rolling under drought stress.
Leaf blade (lamina)
Flat photosynthetic surface of the leaf.
Petiole
Stalk connecting leaf blade to stem.
Leaf margin
Edge of the leaf blade; may be entire, serrated, or lobed.
Entire margin
Smooth-edged leaf.
Lobed margin
Deeply indented edge (e.g., oak, maple).
Serrated margin
Saw-toothed edge.
Sunken stomata
Stomata recessed below epidermis, reducing transpiration.
Glaucous surface
Leaf surface covered with a whitish waxy bloom, reducing radiation load.
Waxy coating
Hydrophobic surface preventing water loss and pathogen entry.
Scale leaves
Small, scale-like leaves (e.g., cedars, junipers).
Simple leaf
Single blade (e.g., oak, maple).
Compound leaf
Blade divided into leaflets (e.g., ash, walnut).
Pinnately compound
Leaflets arranged along a central axis (rachis).
Pinnately compound leaf
Leaflets arranged along both sides of a common axis.
Palmately compound
Leaflets radiating from one point (e.g., horse chestnut).
Palmately compound leaf
Leaflets radiating from a single point (like fingers from a hand).
Oblong leaf
Longer than wide with nearly parallel sides.
Cordate leaf
Heart-shaped leaf.
Ovate leaf
Egg-shaped, broader at base.
Linear leaf
Very narrow and elongated.
Lanceolate leaf
Long, narrow, tapering to tip.
Lobed leaf
Leaf with deep indentations but not divided into leaflets.
Needle leaf
Narrow, needle-like leaves typical of conifers.
Needle leaves
Thin, needle-like leaves (e.g., pines).
Drip tip
Extended pointed tip aiding water runoff in tropical rainforests.
Succulent leaves
Thickened leaves for water storage.
Deciduous leaves
Leaves shed seasonally.
Evergreen leaves
Leaves persisting multiple years.
Photosynthesis equation
6 CO₂ + 6 H₂O + light → C₆H₁₂O₆ + 6 O₂.
Overall photosynthesis
6 CO₂ + 6 H₂O + light → C₆H₁₂O₆ + 6 O₂.
Calvin cycle
In stroma: uses ATP & NADPH to fix CO₂ into sugars.
Light reactions
In thylakoids: convert light → ATP & NADPH, release O₂.
Photorespiration
Rubisco reaction with O₂ reducing photosynthetic efficiency.
Respiration
Breakdown of sugars to release ATP and CO₂.
Diffusion
Passive movement of CO₂ and O₂ through stomata/air spaces.
Osmosis
Water movement across membranes due to solute gradients.
Transpiration
Loss of water vapor through stomata.
Stomatal conductance
Measure of gas exchange capacity through stomata.
Leaf cooling
Temperature regulation via transpiration and convection.
Water Use Efficiency (WUE)
Ratio of photosynthesis (carbon gain) to transpiration (water loss).
Opening (stomata)
K⁺ enters guard cells → water uptake → swelling opens pore.
Closing (stomata)
K⁺ exits guard cells → water leaves → cells shrink, pore closes.
Chlorophyll
Green pigment absorbing red/blue light for photosynthesis.
Carotenoids
Accessory pigments that absorb excess light and provide photoprotection.
Anthocyanins
Pigments that provide red/blue coloration and photoprotection.
Accessory pigments
Carotenoids, xanthophylls that expand light absorption and protect chlorophyll.
PAR
400–700 nm light usable in photosynthesis.
PAR (Photosynthetically Active Radiation)
Light in the 400–700 nm range used for photosynthesis.
Action spectrum
Wavelengths most effective in photosynthesis.
Absorption spectrum
Wavelengths absorbed by pigments.
Photoinhibition
Reduction of photosynthetic capacity due to excess light.
Reflection
Portion of light (especially green) reflected by leaves.
Transmission
Light passing through leaves.
Photons
Packets of light energy; shorter wavelengths = higher energy.
Energy balance equation
Rn = H + LE + G (net radiation = sensible heat + latent heat + ground heat).
Rn (Net radiation)
Absorbed – emitted energy balance.
H (Sensible heat flux)
Heat transfer by conduction/convection.
LE (Latent heat flux)
Energy loss through evaporation/transpiration.
G (Ground heat flux)
Heat stored in or released from leaf/ground.
Light compensation point
Light intensity at which photosynthetic CO₂ uptake equals respiration CO₂ release.
Light saturation point
Light intensity at which photosynthesis no longer increases with additional light.
Parallel venation
Veins run parallel (typical of monocots).
Reticulate venation
Branching vein network (typical of dicots).