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🧮Species richness
The number of species in a community.
🧮Five possible productivity–richness relationships
U-shaped, negative, none, positive and hump-shaped.
🧮Mittelbach et al. 2001
Reviewed many studies and found the hump-shaped productivity–richness relationship was the most common in nearly every group.
🧮Why richness is hump-shaped with productivity
At low productivity few species can survive; at intermediate levels many coexist; at high productivity a few strong competitors take over.
🧮Park Grass Experiment
Started in 1856 at Rothamsted, England. Adding more types of resources (fertilizers) to grassland plots lowered the number of plant species.
🧮Park Grass regression
Species = 34.1 − 3.1 × (number of resources), R² = 0.49, p = 0.02. Each added resource meant about 3 fewer species, and resources explain about half the variation.
🧮MacArthur & MacArthur 1961
Foliage height diversity (number of vertical vegetation layers) predicted bird species diversity better than plant species diversity did.
🧮Why habitat structure increases diversity
More structural layers create more niches, so more species can coexist.
🧮Intermediate Disturbance Hypothesis
Species richness is highest at intermediate (occasional, moderate) disturbance, compared with rare or very frequent disturbance.
🧮IDH: low disturbance
Dominant competitors exclude other species, so richness is low.
🧮IDH: high disturbance
Only a few tough or fast-colonizing species survive, so richness is low.
🧮Lubchenco 1978
🦠Algal species richness in tide pools peaked at intermediate snail (grazer) densities.