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when both species are benefited
mutualism
when only one species is benefitted and the other is not affected
commensalism
when one species is benefitted and the other is harmed
exploitation
when both species are hurt
competition
when one species is not affected and the other is hurt
amensalism
types of exploitation
allelopathy, herbivory, predation, parasitism
competition with members of own species is what type of competiton
intraspecific
competition between individuals of two species (reduces fitness of both) is what type of compeition
interspecific
direct aggressive interaction between individuals observed short term
interference
type of competition where no need for individuals to interact directly and outcome determined by how efficiently each competitor use the resource
resource
interactions in which most of all of the life cycle of one organism occurs inside or on another
symbiosis
when there is a limited supply of suitable space for a species
resource limitation
ecological principle describing how plant mortality occurs in crowded, even-aged monocultures due to intense competition for limited resources like light, water, and space
self-thinning (3/2) rule
first to study competition in 1917
Arthur Tansley
observed closely related species occurring together often grow in different habitat and asked if interspecific competition led to separation of two species on plants on different soil types
arthur tansley
arthur Tansley looked at bedstraw species in what habitats
bedstraw grown on acid soil and alkaline limestone hills
what did Tansley find about Galium grown on the two soil types
they grew on both soil types so plants not limited to a certain soil type
results of only G. saxatile on acid vs. basic soil
results of only G. sylvestre on acid vs. basic soil
results of both species planted together on acid vs. basic soil
did better on acidic soil / did better on basic soil / the basic adapted G. sylvestre grew better than the G. saxatile on basic soil and vice versa but both grew
Tansley discovered that species were adapted to (__), which is the ecological space where species are specialized upon. There was a clear (__) of native species in soil they are adapted for
realized niche / competitive dominance
first attempted to model interspecific competition was (__), based models on (__) equation which is a model of (__) competition
lotka and volterra / logistic / intraspecific
in the logistic equation, the term N/K measures
effect on growth of population by adding new member of same species (intraspecific competition)
what is the modified logistic growth formula for interspecific competition by lotka and volterra
dN/dt = r1N1 ([K1-N1-α12N2]/K1)
proposes that two species with identical niches cannot coexist indefinitely, which leads to the prediction that coexisting species will have different niches
competitive exclusion principle
since the effect of one species on the growth of another would not likely be identical to a species effect on its own growth, (__) must be made to analyze both inter and intra specific competition
competition coefficient
alpha, competitive effects of competing species
competition coefficent
what does α21 mean
effect of species 1 on species 2
what does α12 > 1 mean
competitive effect of species 2 on species 1 is greater than effect of individual of species 1
based on lotka and volterra equation, when do we know populations of both species stop growing
when dN/dt equals 0
equations for straight lines where everywhere along the lines population growth is stopped
isoclines of zero population growth
above an isocline of zero growth, the population of a species is (__). below it, the population is (__)
decreasing / increasing
what happens when N1= K1
environment is completely filled by species 1 and no species 2
the Lotka-Volterra model predicts that one species will exclude the other when the isoclines
do not cross
if the isocline for species 1 lies above that of species 2, species 1 will
eventually exclude species 2
coexistence between species is only possible when the isoclines
cross
the lotka-volterra model predicts that species coexist when…
intraspecific competition is stronger than interspecific competition

at K1, species (__) wins, population size equals (__). (__) strength is stronger for species (__)
1 / K1 / interspecific / 1

at K2, species (__) wins and population size equals (__). (__) strength for species (__) is stronger
2 / K2 / interspecific / 2

there is (__) strength when they cross. eventually, (__) wins. coexistence eventually drives one species (__)
equivalent / species 1 or 2 / extinct
how many outcomes of competition by lotka and volterra are there
4
Gause used (__) to model Lotka-Volterra competition
P. aurelia and P. caudatum paramecium
studied would one of these two species drive the other to extinction if grown together in microcosms where they were forced to compete with each other for a limited food supply
Gause
difference between P. aurelia and P.caudatum
P. aurelia is smaller, has 2 micronuclei and posterior end is more round whereas P.caudatum has cylinder cigar shape and one micronucleus
Gause demonstrated resource (__) by growing pure populations in presence of two different food concentrations. He observed (___) growth with obvious (__) at both full and half strength concentrations of food. when food availability was halved, the carry capacity was (__). thus when grown alone, carrying capacity was determined by
limitation / sigmoidal / carrying capacities / reduced / intraspecific competition
when grown together, what did Gause find
P. aurelia survived but P. caudatum declined
what did Thomas Park study to show outcome of competition depends on environment, they infest stored grain products
flour beetles
Thomas Park found that in hot-wet environments, species of flour beetles grow separately had what growth? what about when grown together?
both species had healthy and persistent populations / only one species sustained
under such intermediate conditions, the species establishing itself in greater numbers first generally wins out in competition, a phenomenon known as
priority effect
with flour beetle experiment, outcome (___) assured because (__) conditions and winner was (__)
not / intermediate / unpredictable
if the competitive exclusion principle is in place, how do species coexist?
natural selection reduces competitive pressure by specialization into realized niche
what did Robert MacArthur show with warbler species about competition
resource partitioning. warblers coexist on same resource by subdividing niche
competition can have short term ecological effects on niches of species by restricting them to (_) yet these species may retain their capacity to inhabit the fuller range of environments called (__)
realized niche / fundamental niche
t or f: if competitive interactions are strong and pervasive enough, they may produce an evolutionary response in competitor population, an evolutionary response that changes fundamental niche
t
Connell discovered that (__)plays a key role in determining the lower limit of Chthamalus barnacles within the intertidal zone
interspecific competition
interspecific competition can lead to (__), the circumstance in which two species differ more from each other in geographic areas where they occur together than wehre their distributions do not overlap
character displacement
resource partitioning and niche specialization caused by competition leads to
niche differentiation
ecological space where competition is most severe where two species occupy mutually exclusive microhabitats in nature
niche overlap
niche overlap due to combination of
competition and difference in fundamental niche
two species living in separate geographical ranges
allopatric
two species occuring together
sympatric
when species are allopatric, their beak sizes are (__), but beak sizes are (__) when species are sympatric
similar / different
what was found on Santa Maria, San Cristobal where G. fuliginosa and G. fortis coexisted?
what was found on Los Hermanos where G. fuliginosa was allopatric?
what was found on Daphne where G. fortis was allopatric?
no overlap between beak sizes / its beak size could get larger because no competition from other species / its beak size could be smaller because no competition from other species
what do emerald jewel wasps do to cockroach and what type of relationship is this
inject pasasitoid venom that paralyzes cockroach and lays egg on roach. larvae burrow into cockroach and feed off of it / exploitation
populations of a wide variety of predators and prey are not (__) but rather (__) in abundance over periods of days to decades. increases/decreases between predator and prey (__) each other and fluctuations are (__). (__) always cycles higher
static / fluctuating dynamically / mirror / regular / prey
Lotka Volterra assumes host population grows (__) and population size is limited by (__)
exponentially / predators
explain the meaning of lotka-volterra host population growth equation
dNh/dt = rhNh - pNhNp
rate of host population change equals exponential rate of increase by host population (per capita rate of increase times number of hosts) minus number of prey or hosts killed by predator (number of hosts times predation rate times number of predators)
explain the meaning of lotka-volterra predator population growth equation
dNp/dt = cpNhNp - dpNp
rate of predator population change equals rate at which hosts are converted to predator offspring (host to predator conversion rate) minus number of predator deaths
a rise in prey population is followed by (__) in predation. an increased number of predators causes a (__) in prey population. as prey population falls, it will support (__) predators. with fewer predators, the prey population (__) again. this create a cycle of (__)
rise / decrease / less / rises / oscillations
during Gause’s experiments with paramecium and protist didinium, in the absence of refuges and immigration, what happened to the populations.
predator quickly consumed all paramecium and both went extinct
during Gause’s experiments with paramecium and didinium, adding a refuge like (__) affected prey how? predator how?
sediment / gave paramecium somewhere to hide so population persisted / predator became extinct
during Gause’s experiments with paramecium and didinium, cycles maintained only when simulated (__) of (__)
immigration / predator
situations in which members of an exploited population have some protection from predators and parasites
refuges
was Huffaker able to see the oscillations in his experiment on mites and oranges
only after making it complex and using 120 oranges
how does complexity of environment affect stability and interactions and species
complexity increases stability and helps maintain interactions and number of species
how does availability of refuges affect p in Lotka equation
p (rate of predation) is lower when more refuges
t or f: numerical response eventually levels offa t the point where further increases in prey density no longer produce increased predator density
t
occasional protection in large numbers over short time that prevents predators from building up
predator satiation
list the variety of refuges utilized by prey
spatial, flight, size, large groups, habitat, behavioral
list an example of crypsis
moth resembles dead leaf on forest floor in Costa Rica
(__) store two chemicals (hydroquinone and hydrogen peroxide). when threatened, the two chemicals are (__) and undergo violent reaction. boiling foul smelling liquid becomes a (__) and expelled with popping sound. this is an example of (__) defense
bombardier beetles / mixed / gas / chemical
list ways prey avoid predators
refuges, crypsis, chemical defense, mechanical defense, mimicry, behavior
poison arrow frogs utilize (__), which is an example of (__) defense. they have a brilliant advertisment of toxicity. golden dart frog native to columbia currently considered most poisonous vertebrate also utilizes this.
aposematic coloring / chemical
example of mechanical defense
porcupine needles, turtle shell
what makes up the mimicry complex and explain parts
model (dangerous organism that is copied), mimic (nondangerous organism copying model), and dupe (predator being fooled)
list an example of batesian mimicry
caterpillar backside looks like venomous snake head or non-venomous snake looks like venomous snake
how do two snake populations have to live for batesian mimicry to occur
sympatric
playing dead is a behavior used to avoid being eaten by predators because many predators like fresh meat. this is called
catalepsis
the eastern hognose snake uses (__) to avoid predators. they (__) necks and (__) loudly. if that doesn’t work, they (__) by rolling over with tongue out and remain limp. if that doesn’t work, they may (__)
intimidation / inflate / hiss / play dead / bite
synchronous widespread seed and fruit production is known as
masting
how do oak trees use predator satiation (swamping)
produce many acorn fruit only occasionally so cicadas cannot specialized on resource
relationship that increases fitness of one species while reducing fitness of other
parasitism
(__) consume tissues or body fluids of organism on which it lives called the (__). they typically harm but do not (__) the organism they eat
parasites / host / kill
t or f: parasites are found in all kingdoms
t
bacteria, fungi, and protists, usually microscopic organisms that cause diease
pathogen
number of host species that a single species of parasite can infect
host range
do parasites have a large of small amount of possible host species? this is called?
small / host specificity
host range limits due to (__) processes that evolve in (__) with hosts. as frequency of parasite trait increases, it will select for gene (__) to parasite in host. (__) is always a step ahead and (__) is trying to keep up
coevolutionary / parallel / resistant / parasite / host
species must constantly adapt and evolve against co-evolving opposing species just to survive and maintain their relative fitness is called the
red queen hypothesis
when you remove the ability to keep changing and evolving, such as if a population gets small and loses diversity, the population will be pushed to
extinction
for an evolutionary system, continuing (__) is needed to maintain fitness. why are cavendish bananas at risk of extinction?
development / the banana is triploid and cannot sexually reproduce so more susceptible to Panama Disease since no diversity
list an example of parasites altering host behavior
worm goes into arthropod. usually arthropod resides at bottom on pond, but the worm causes it to go up towards the light, leading to a higher chance of being eaten by a duck and allows it to produce more worms