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ecotherm
regulate body temp via energy exchange with the enviroment
cold blooded
ex. sunlight, warm surfaces
they lack baseline "leaky" cellular burn and have far fewer mitochondria, so their cells simply don't generate enough heat at rest.
Even if an ____ could suddenly boost its metabolic heat output, that heat would instantly radiate away into the air or water.
digestive system and metabolic pathway are optimized to operate on very few calories. They cannot process food fast enough to supply the constant fuel required for endothermy.
endotherm
rely on internal heat generation (mostly birds and mammels)
warm blooded
ex. shivering, sweating
cells are packed with mitochondria, and their cell membranes are intentionally "leaky" to ions . Pumping these ions back and forth constantly burns ATP, which generates continuous internal heat.
enzymes are evolutionary fine-tuned to function within a very narrow, high temperature range. If an ____ allows its body temperature to drop to match a cold environment (hypothermia), its cellular enzymes slow down dramatically, leading to loss of consciousness and death
basal metabolic rate (BMR)
minimum energy (calories) an organism needs to stay alive at complete rest
problem with small aquatic ectotherms
metabolic heat made by swimming instantly loses internal energy, resulting in being in the same temp as water
regional endothermy/ mesothermic
trick where ectotherm keeps warmer than enviroment
tuna keep muscles contracted and use special netowrk of bloodvessels to keep blood warmed and cycling away from spine, warming new cold blood
terristerial ectotherms ____ to adjust temp
move around
insects and reptiles stay safe using
camoflauge
avoidance of the cold in ectotherm include
seasonal migration
microsites
main goal is to avoid damage from ice formation in cells
glycerol helps
lower freezing point of bodily fluids
keeps ice crystals outside cell
vertebraes do not tolerate freezing because
ice crystals physically damage cells
result in severe osmotic dehydration
blocked circulation
In animals that withstand freezing, the freezing water is limited to _____
the space outside the cells
ice nucleating proteins outside the cell controls ice formation
glucose and glycerol are made inside the cell to lower freezing point
The cost of being endothermic is a high demand for
energy (food) to support metabolic heat production
Thermoneutral zone
range of environmental temperature that a constant BMR can be maintained
The range of outside environmental temperatures where an endotherm maintains its normal body temperature
Inside this temperature window, the body doesn't spend extra metabolic energy shivering or sweating to stay regulated
lower critical temperature
when heat loss is greater than metabolic production ; body temp drops and heat generation increases
temp below the lowest temp body can handle
Because heat is escaping faster than the baseline engine can generate it, the body must ramp up active heat production
The colder it gets below the LCT, the higher the metabolic rate must go to prevent core body temperature from dropping
incipient species
a population of organisms in the early, ongoing process of speciation where they begin to diverge genetically but can still occasionally interbreed
autotroph
assimilate radiant energy from sunlight or inorganic compounds
energy is converted into chemical energy stored in bonds of organic molecules
ex. photosynthesis, chemosynthesis
heterotroph
obtain energy by consuming organic compounds from other organisms
some consume non-living matter
parasites and herbivores consume live hosts but don’t kill them
predators capture and consume live prey
holoparasites
plants that are non-photosynthetic
get energy from other plants
dodder
holoparasite that’s an agricultural pest and can significantly reduce biomass in host plant
hemiparasite
photosynthetic plant but still obtains nutrients
ex. mistletoe
photosynthesis
sunlight provides energy to take up co2 and synthesis organic compounds
light reactions (light absorptions) + dark reactions (co2 is fixed and uses light energy to synthesize carbs)
most common source of energy on earth
photosynthetic rate
determines supply of energy which influences growth and reproduction
light response curve
show influence of light levels on photosynthetic rate
light compensation point
where co2 uptake is balanced by co2 loss by respiration
photosynthesis (co2 uptake) = cellular resp (co2 loss)
saturation point
when photosynthesis no longer increases even when more light is absorbed
chemosynthesis
energy from inorganic compounds is used to produce carbs
important in nutrient cycling bacteria and some ecosystems like hydrothermal vent communities
autotrophic process
specialized bacteria and archaea build organic food from carbon dioxide using energy harvested from inorganic chemicals instead of sunlight
primary pigment for light harvesting
chlorophyll
plants can acclimatize to
changing light intensities
In high light, a plant builds thick, dense leaves packed with extra enzymes (like Rubisco) and photoprotective pigments, allowing it to process intense energy quickly without overheating
in low light, the plant lowers its baseline energy bill by reducing excess enzymes, building broad, paper-thin leaves that maximize surface area to catch every stray photon, thereby dropping its light compensation point
shifts in light saturation point involve
morphological and physiological changes
a plant's physical structure and enzymatic machinery dictate its maximum operational capacity
result of matching its metabolic investment to the light availability of its environment
adjust to keep up with more or less light
photorespiration
Photorespiration is a wasteful metabolic pathway that occurs in plants when the enzyme Rubisco binds to oxygen instead of carbon dioxide
rubisco catalyzes two competing reactions
carboxylase reaction
oxygenase reaction (photorespiration)
benefits of photorespiration
photorespiration may protect plants from damage at high light levels
When plants absorb excess light energy or close their stomata during drought, the electron transport chain can become dangerously overloaded; photorespiration acts as a crucial "energy safety valve" by consuming surplus ATP and NADPH, preventing the accumulation of reactive oxygen species (ROS)
Photorespiration is a large disadvantage if co2 is low and temp high
C4 PHOTOSYNTHETIC PATHWAY
evolutionary adaptation that prevents photorespiration by physically separating initial carbon capture from the Calvin cycle
outside CO2 enters the mesophyll cells (outer layer).
An enzyme called PEP carboxylase fixes onto phosphoenolpyruvate (PEP) to form a 4-carbon compound (oxaloacetate/malate)
PEP carboxylase has zero affinity for oxygen, so it never makes the mistake Rubisco does, even when CO2 is low
PEP carboxylase is broken down into CO2
Rubisco fixes the CO2 under the normal Calvin cycle with virtually zero photorespiration
CRASSULACEON ACID METABOLISM (CAM)
alternative photosynthetic adaptation that minimizes photorespiration and prevents water loss by separating carbon capture and the Calvin cycle by time (night vs. day)
at night, stomata opens and co2 fixations occurs by PEP
during the day stomata is sealed to prevent water loss
factors affecting ecological success (enviromental)
extreme conditions that exceed tolerance limits(stress)
availibility of energy (affects growth and reproduction)
energy supply (influences tolerance of the environment)
climate envelope
set of climate conditions—such as temperature ranges, precipitation levels, and seasonal patterns—under which a particular species can survive, reproduce, and maintain a stable population
physiological ecology
study of environmental influence on survival
Processes have optimal conditions for function
stress
deviations from optimal conditions
lead to lower rates of physiological process and decreased growth and survival
acclimatization
Adjusting to stress through behaviour or physiology
ussually short term and reversible
ex. climbing mount everest takes a few days to acclimatize to changing elevations
adaptation
resulted from natural selection
coping with stress leads to natural selection, favoured traits become more common leading to adaptation
response to stress
speciation
when species are separated for so long that that natural selection results in them becoming reproductively isolated, become different species
ecotypes
populations with adaptations to unique environments
Ecotypes can eventually become separate species as populations diverge and become reproductively isolated
Members of different ecotypes are still the exact same species—they can interbreed and produce fertile offspring—but they look, grow, or function differently because natural selection has tweaked their DNA to survive local conditions
Survival and functioning of organisms is strongly tied to their
internal temperature
isozymes
special enzymes produced by some species that have different optimal temperatures, allows for acclimation to changing conditions
SA:V
the smaller the ectotherm, the less energy it can hold but the faster it can store and lose heat
large ectotherms can hold heat for longer
large ectotherms adjust heat by
-vasodialtion/vasoconstriction (skipjack tuna)
small ectotherms adjust by
changing locations
changing colour of skin
ectotherms tolerate/avoid freezing by
seasonal migration
relocate to microsites (underground burrows)
microsite
underground burrows that are above freezing temps
Cryonics
the preservation of bodies by freezing, in hope that they can be brought back to life in the future
The cost of being endothermic
a high demand for energy (food) to support metabolic heat production
Metabolic rates are a function of
the external temperature and rate of heat loss
represents the speed at which an organism burns chemical energy (ATP) to power cellular work.
chemical reactions are sensitive to heat and heat constantly flows down thermal gradients, metabolic rate is fundamentally tied to both external temperature and the rate of heat loss.
dormancy
state where little to no metabolic activity occurs
method of escaping extreme heat or cold
evolution of endothermy required
insulation, like fat, fur and feathers
Mammals in the Arctic have ___ critical temp than tropical ones
lower
can withstand colder temps without needing extra atp
The rate of metabolic activity increases more rapidly below the lower critical temperature in
tropical mammals as compared to Arctic mammals.
tropical mamels have a higher LCT
metabolic activity increases to tolerate freezing temps
Metabolic rate
how fast your body burns fuel (calories) to generate energy (ATP)
torpor
state of dormancy observed in mammals and birds
Body temperature and basal metabolic rates are low, which conserves energy
methods to tolerate heat
spray water
open mouth
moving to shade
evaporative heat loss (sweating, licking), wicks away heat in form of water to gas
plants adapt to heat via
transpiration
water that’s absorbed by plant roots evaporates through stomata
if soil is limited
too much transpiration results in water loss, stomata closes but then overheating may occur
plants may shed their leaves during dry season to reduce sa that requires water
pubescence
hairs on leaf surfaces that reflect solar energy
lowers the total heat absorbed by the plant, keeping leaf temperatures cooler under intense sunlight
convection
transfer of heat from solid to air (traps layer of warm air)
wind destroys
high altitude alpine plants avoid convection loss by
staying close to ground to avoid high winds
some have insulating hair
some organisms tolerate arid conditions by
going into suspended animation (stop all metabolic processes)
torpor vs suspended animation
torpor still maintains low levels of metabolic process, thus require high fat stores
suspended animation consumes no atp and no reactions
reptiles are very successful in dry conditions because
They have thick skin with layers of dead cells, fatty coatings, and plates or scales
skin is impermeable barrier against water loss
fatty compounds embedded in their skin layers repel water, creating an airtight seal against evaporation
sweat glands in mammels are a tradeoff between
water loss resistance and evaporative cooling
their skin is sealed, so reptiles cannot sweat. To cool down, they rely entirely on behavior (seeking shade, burrowing) or minor evaporative cooling through their mouths (gaping)
in mammals, Sweating releases liquid water directly onto the skin, providing unmatched evaporative cooling during intense heat or high physical activity, but It requires burning through large amounts of body water. In a desert environment where liquid is scarce, relying heavily on sweating can trigger lethal dehydration and osmotic collapse very quickly
ecology
study of interactions between organisms and their enviroment
the distribution, locations and abundance of organisms
how systems adjust and change during disruptioons
species ____ play equal roles in maintaining fn of ecosystem
DON’T
hierachar of ecosystem
organism , population, community, ecosystem, biosphere
organism
individual
population
species living together at the same time, can interbreed
community
population of different species
ecosystem
biotic + abiotic in an enviroment
biosphere
all ecosystems on earth
producer
organism that uses energy from external source to produce energy
consumer
obtains energy by eating other organisms or organic matter
net primary production (NPP)
amount of energy per unit of time that producers fix by photosynthesis or other means - amount used in cellular respiration
nutrient cycle
cyclic movmeent of nutrient between organism and enviroment
cellular respiration
biochemical process by which cells break down nutrient molecules (primarily glucose) to harvest energy in the form of adenosine triphosphate (ATP)
virtually ALL living organisms perform .
Multicellular animals also have evolved specialized
tissues and organs for absorption, digestion, transport, and excretion
Compared with omnivorous humans, herbivorous primates have longer
digestive systems
Heterotrophs have tremendous diversity in morphological and physiological
feeding adaptations
Plants that thrive at low temperatures have higher proportions of
unsaturated lipids (with double bonds) in their cell membranes
evolution is
allele frequency change
Takes millions of years
Due to the environmental pressures, the frequency of specific alleles changes
change in organisms over time
directional selection
individuals of one phenotype extreme is favoured
ex. large ram horn size favoured, over time the average ram horn size increases
Stabilizing phenotype
intermediate phenotype favoured
ex. human baby birth weight
disruptive selection
both extremes of phenotype favoured
genetic drift
evolution caused by random events (affects size of gene pool)
significant/damaging in small populations
as alleles go missing, genetic variety lessens
gene flow
transfer of alleles between separate populations
increases diversity within the population but decreases differences between the different groups
constrains on evolution
Lack of genetic variation (natural selection can only choose from existing alleles or mutations, otherwise may be at risk of extinction if adaptation cannot occur. limits adaptation. adaptation cannot occur if beneficial allele doesn’t exist)
Evolutionary history (organisms have physiological rules that dictate how their body can grow and change, can only adopt certain traits)
Ecological trade-offs (every trait comes with pros and cons, must consider if energetic costs outweigh benefits)
keystone species
help define an ecosystem, has large effect on ecosystem
controls community structure and biodiversity
large-scale processes that drive evolution
speciation
adaptive radiation
extinction
evolution vs adaptation
Evolution is the overall change in the genetic composition of a population over successive generations, whereas an adaptation is a specific inherited trait or mechanism that enhances an organism's ability to survive and reproduce in its environment
Adaptation is primarily caused by natural selection, whereas evolution is also caused by genetic drift, gene flow , mutations, and is more about change in allele frequency rather than adaptation to the environment
speciation
the process by which one ancestral lineage splits into two or more distinct, genetically independent species
when gene flow between two populations stops, they evolve independently and eventually become reproductively isolated
adaptive radiation
when a single ancestral species rapidly evolves into a wide variety of descendant species, each adapted to a specific ecological niche or lifestyle
when adaption occurs across many species as they adapt to a new enviroment
Extinction
Loss of species in an ecosystem
Extinctions occur all the time, only a few huge mass extinction events occur
show patterns of evolutionary change
mass extinctions can increase diversity by removing competitors and allowing survivors to rise to new species
Three types of natural selection
1. Directional selection: Individuals at one phenotypic
extreme (e.g., large size) are favored.
2. Stabilizing selection: Individuals with an intermediate
phenotype are favored.
3. Disruptive selection: Individuals at both phenotypic
extremes are favored
Evolution is viewed as
genetic change (allele frequencies), over time or as a process of descent with modification