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What Is Biodiversity?
The variety of all living organisms on Earth.
what does biodiversity include
includes all forms of life, visible and microscopic, and the complex web of interactions between them and their environments.
3 types of bio diversity
Genetic diversity
Species diversity
Ecosystem diversity
Genetic diversity
Genetic differences within a species
Species diversity
Diversity among species within an ecosystem.
Ecosystem diversity
Variety of ecosystems, such as forests and wetlands.
Species richness
the number of different species in an area.
formal species descriptions
Scientists identify, classify, and officially describe species.
Undiscovered species
Many species exist that scientists have not formally described.
Estimated global species richness
Scientists estimate how many species exist worldwide, including species not yet discovered.
DNA identification
DNA can reveal that organisms that look very similar actually belong to different species.
Species abundance
How many individuals of each species are present.
Species evenness
How evenly individuals are distributed among the species.
species-area relationship
Larger areas generally contain more species.
Alpha diversity
diversity of species in a single habitat
Beta diversity
difference in species composition between two or more habitats
Gamma diversity
Total species diversity across a large region
Species Rank Curves
A steeper curve indicates greater differences in abundance from the most common to the least common species.
A longer line indicates higher species richness.
Seven Forms of Rarity-Geographic range
How large or small an area the species occupies.
Seven Forms of Rarity-Habitat specificity
Whether it can live in many habitats or only a narrow range of habitats.
Seven Forms of Rarity-Local population size
How many individuals are present locally.
Widespread, Common rarity
geographic range - large
habitat specificity - Broad
local population size - large
Widespread, Rare
geographic range - large
habitat specificity - narrow
local population size - small
Restricted, Common
geographic range - small
habitat specificity - broad
local population size - large
Restricted, Rare
geographic range - small
habitat specificity - narrow
local population size - small
Frequent Type Rarity
Geographic range - large
Habitat specificity - narrow
Local population size- sparse
Endemic, Common
geographic range - very small
habitat specificity - broad
local population size - large
Endemic, Rare
geographic range - very small
habitat specificity - narrow
local population size - small
Functional Diversity Dendrograms
Maps the differences between
functional traits among species.
• Usually feeding niche
Sum total of branch lengths is a
measure of community functional
diversity
Functional Diversity
Larger circles = larger FD
Warmer colors = greater FD loss

Phylogenetic Diversity
A phylogeny of all known lemur taxa (blue branches)
The phylogeny can identify species that are unique.
Global geographical patterns in biodiversity
Tropical regions generally have the highest species richness.
intertidal zone
Transitional coastal regions
Alternately exposed and submerged by tides
High diversity
Zonation
is a vertical banding of the
organisms living on the rocky coastline.
Spray or Splash Zone
uppermost area exposed mainly to sea spray
high tide zone
area covered by water mainly during high tides and exposed to air for much of day
middle tide zone
area alternately exposed and submerged as tides change
low tide zone
area submerged most of the time, exposed mainly during high tides
Biotic factors affecting organisms living in the intertidal zone:
living influences such as competition, predation, and reproduction
Abiotic factors affecting organisms living in the intertidal zone:
nonliving influences such as temperature, salinity, waves, and dissolved oxygen
stress-waveshock
Adaptation, Example
Adherent threads: Mussels
Stick tightly: Chiton limpets
Conical shape: Barnacles, limpets
Flexible stalks: Kelp, Gooseneck Barnacle
Motility & hide in cracks: Crabs, Periwinkles
stress-dehydration
Adaptation Example
Motility; Sea stars, Periwinkles
Operculum; Periwinkles, Turban snails
stick tightly; chiton, limpets
Seal openings; Barnacles, mussels
stress-temperature extremes
Motility: Crabs
Evaporative cooling: Mussels
Tolerance: Barnacles
Nocturnal behavior: Rock Lice
stress-Salinity extremes
Tolerance, Periwinkles
Motility, Crabs
stress-Feeding
Suspension feeding: Mussels, barnacles
Grazing: Chiton, limpets, sea urchins
Detritus feeding: Crabs, sea hares, some snails
Predation: Octopus, sea stars, some snails and nudibranchs
Absorption of DOM: Sea urchins
stress-Competition
Aggression: Octopus, crabs
Rapid growth: Barnacles
Opportunistic larval establishment: Mussels, barnacles
Large number of offspring: Sea urchins, barnacles
Good dispersal: Species with planktonic larvae
stress-predation
Hard Shells: Barnacles, mussels
Motility: crabs
Toxins: Nudibranchs, kelps
Protective Coloration: Sculpins
Antipredation behaviors: Limpets
The Keystone Species of the Intertidal Zone
Pisaster (Sea Star) defends the rocky intertidal zone from being taken over by mussels, barnacles.
Primary Producers of the Intertidal Zone
Devil’s Bath Towel – Gigartina sp.
A type of red seaweed
Uses sunlight to make food through photosynthesis and provide habitat for other organisms.
Aplysia californica
releases a purple fluid that can blind predators. The fluid is synthesized from red algae they graze on
What is competition?
The interaction between organisms or species that compete for limited resources, such as food, water, or space, which can negatively affect their fitness.
Interspecific competition
competition between different species
Intraspecific competition
competition within species of same type
An ecological niche
is the role and position a species has in its environment, including how it meets the needs for food, shelter, and reproduction, as well as its interactions with other species.
what does ecological niche encompass
It encompasses all biotic and abiotic factors necessary
for survival and reproduction.
Competitive Exclusion Principle
No two species can occupy the same niche at the same time in the same environment.
If two species overlap substantially…
• One will be outcompeted and driven to extinction.
• Natural selection will favor individuals of both species that avoid
competition
resource partitioning
Reduced competition among coexisting species as a result of differing niches
Resource partitioning may include
feeding, nest sites, and other aspects of a species’ ecological niche
the fundamental niche
The potential niche of a species. Various factors, such as competition, may exclude a species from its fundamental niche
The realized niche.
The lifestyle a species actually pursues and the resources it actually uses
Character Displacement
The divergence in traits in two similar species living in the same area
The Lotka-Volterra Model
1.Used to estimate the populations of predator-prey systems
Used to estimate populations of competitors
carrying capacity (K)
is the maximum population density an environment can sustain.
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Stable coexistence
both species persist; their populations return toward a stable equilibrium.
Unstable coexistence
the winner depends on initial population densities; the system moves away from the equilibrium.
Zero Net Growth Isoclines and R
The amount of two
resources of interest
to a species are on
each axis.
Rx*
is the threshold level of resource X at which births and deaths are balanced….Hence Zero Net Growth. B = D
The resource supply point (S)
represents the natural availability of the resources in the environment.
The green resource vectors
always point toward the RSP. Consumption vectors point in the opposite direction.
Rx*, Ry*
the point where B = D; the critical resource levels
Exploitation
the consuming of one species by another, is a common and important force in nature. A simple model of exploitation is a (+, -) interaction
There are three main types of interaction
• Predator – Prey
• Parasite – Host
• Herbivore – Plant
Exploiter
(predator) gets a fitness benefit from consuming prey.
Prey
experience fitness loss because they have been eaten or partially consumed.
Detritivory
consuming dead prey is NOT predation
Possible Outcomes of Exploitation
1. Extinction of the prey, followed by extinction of the
predator
2. Extinction of predator, followed by increases in the prey
3. Cyclical population dynamics for the exploiter and prey
4. Stable coexistence between exploiter and prey
The Prey Equation
Prey capture rates vary widely
A whale shark might eat million of planktonic organisms
A python might eat a large meal 2-3 times per year
The Exploiter Equation
A predator consuming a prey organism does not translate directly to reproductive output. It takes many prey and several months for a fox to produce a litter of kits.
equilibrium
a condition where population growth is zero, so the population remains constant
zero-growth isocline
line on graph showing combinations of population densities where one population has zero net growth
Density dependence
population growth is affected by population density; prey self-regulation is added through carrying capacity
functional response
how an exploiters rate of prey consumption changes with prey density
allee effect
a population does worse at a low density; a lone buffalo may be more vulnerable to predators than a herd
coevolution
evolutionary changes in one interacting species influence selection on the other species
crypsis
camouflage that makes an organism harder to detect
aposematic coloration
warning coloration that signals a defensive or harmful trait
Batesian Mimicry
A harmless species evolves to mimic a dangerous species
mullerian mimicry
multiple harmful or defended species share similar warning signals
Invasive Species
Non-native and causes ecological harm.
functional response type I
consumption increases roughly linearly with prey density
the slides associate it with neutral stability and cycles
functional response type II
Consumption increases but levels off as the predator becomes limited in how much it can consume.
The slides say it decreases stability, potentially causing extinction.
functional response type III
An S-shaped curve: consumption rises slowly at first, then faster, and eventually levels off.
An efficient exploiter can produce greater stability and damped oscillations.
Evolutionary defenses-predator crypsis
predator camouflage
Evolutionary defenses-Prey crypsis
Zebra stripes providing camouflage in certain surroundings
Evolutionary defenses-Eyespots
Moth caterpillar or moth wings that startle or deter predators