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key hardwood forests in Ontario
- maple, yellow birch, red oak
- beech, white ash, hemlock
- basswood
- black cherry
harvest wood volume
maple (73% of area, 75% of volume), yellow birch, hemlock, beech, oak, ash, basswood, others
plasticity
ability to adapt and cope with environmental changes such as the ability to respond to release
sugar maple silvics
habitat: wide range, even or uneven, pure or mixed stands
shade tolerance: very shade tolerant
establishment: reproduces easily on undisturbed forest floor
development: determined height growth
american beech silvics
habitat: often mixed stand, occasionally pure
shade tolerance: very shade tolerant
establishment: easily regenerates on understory
development: determined height growth
yellow birch
habitat: pure and even aged fire origin stands or mixed stands
shade tolerance: intermediate shade tolerant
establishment: scarification aids regen, wind dispersed small seeds
development: indetermined height growth
red oak
habitat: dry sites
shade tolerance: intermediate shade tolerant
establishment: fire and clearcut best seed bed, responds well to release from moderate suppression
development: determined height growth
basswood
habitat: fast potential growth rate among hardwood species
shade tolerance: seedlings intermediate, mature shade tolerant
establishment: prolific stump sprouter, even age coppice in stand after disturbance, can regenerate entirely vegetatively
development: indetermined height growth
white ash
habitat: considered soil improvement species because high calcium leaf litter
shade tolerance: intermediate shade tolerant, seedlings shade tolerant but tolerance decreases with age
establishment: good response to release, leaf out later than associated broadleaf species. strong apical dominance, low epicomical branching
development: determinate height growth
black cherry
habitat: low plasticity, poor response to release
shade tolerance: shade intolerant, cannot survive under closed canopy
establishment: good natural regeneration, large seeds dispersed by birds
development: determined height growth
assessing individual tree vigor potential
crown position
crown size, leaf area
crown quality
bark character
degree of competition
stand relationships, whole stand view
assessing individual tree quality potential
DBH, larger higher value
site region
cull factor
tree growth rate
cutting cycle
a 20 year cycle to remove basal area, where every cutting cycle removes about 20% basal area
stand stocking
amount of trees in an area related to management objectives. density, crown closure, basal area
key differences between hardwood and softwood forests
phenology: timing of recurring natural evens like budburst, leaf expansions, flowering. boreal forests don’t lose their leaves in the winter
diversity: conifer forests significantly less diverse
vegetative reproduction: most important difference, hardwoods like aspens can reproduce vegetatively through sprouts and suckers
growth and mortality
most critical processes determining the fate of juvenile trees during natural regeneration
competition
an interaction between individual plants that reduces fitness of one or both involved
exploitation
when plants compete by reducing the availability of shared, limiting resources such as light, water and nutrients
interference
when fitness is reduced by mechanisms which do not directly involve limiting shared resources (allelopathy)
4 effects of competition
mutualism, mutually deleterious interaction, asymmetric facilitation, asymmetric competition
competition theories
ruderal: high reproductive and growth rate on disturbed sites
stress tolerator: low reproductive and growth rate on undisturbed sites
competitor: low reproductive effort and high growth rate, productive sites
grime’s theory
assumes that competition intensity increases with site productivity and competition success is defined in the capacity to capture resources
CSR triangle theory
C: high productive/high competition habitats
S: high environmental stress habitats
R: high disturbance (ruderal, open disturbed) habitats
tilman’s theory
total competition intensity is unaffected by habitat productivity though the ratio of aboveground and belowground competition varies. competition success is the ability to draw and tolerate resources to a low level
major mixed forests in BC
red alder, bigleaf maple, doug fir, cedar
paper birch, doug fir, spruce
aspen, paper birch, spruce
65.5% mixedwood in BC
why mixedwood
ecological sustainability: biodiversity, forest health - root rot, insect outbreaks, long term site productivity
multiple values and economic consideration
regulation - forest practice codes
spey creek project thinning
increased understory light
foliar responses
physiological responses
crown architecture
Ht, DBH and volume growth
raft creek project conclusions
thinning overstory birch to 600 spha provides enough light 60% for understory conifers
this density significantly reduced understory vegetation and birch sprouting
birch population research conclusions
birch populations from arid habitats tend to be less productive than populations from less restricted environments
paper birch population studies are better adapted to drought than low soil N
photo induction
there is lag time before a leaf reaches its Amax when exposed to high light after being in low light for a long time
other conclusions
shading to 60% of full light did not significantly reduce growth of all species studied
birch was more productive in mixture
silvicultural policies must be driven less by mill requirements and more by ecosystem dynamics
sugar maple development
determined
sugar maple moisture regime
fresh fertile well drained upland sites
american beech shade tolerance
very shade tolerant
american beech development
determined
american beech moisture regime
moist, fertile well drained fresh soils
yellow birch shade tolerance
intermediate
yellow birch development
indetermined
yellow birch moisture regime
moist, lower slopes, tolerant to poor drainage
red oak shade tolerance
intermediate
red oak development
determined
red oak moisture regime
dry sites, fresh to moist, well drained
basswood shade tolerance
seedlings intermediate, mature shade tolerant
basswood development
indetermined
basswood moisture regime
fresh to wet, moderately rich ecosites
white ash shade tolerance
intermediate
white ash development
determined
white ash moisture regime
very fertile, fresh to moist, well drained sites
black cherry shade tolerance
intolerant
black cherry development
determined
black cherry moisture regime
cool, moist, well drained sites
key differences between hardwood and softwood forests
phenology, diversity, vegetative reproduction
shade tolerance
the ability of trees to survive and maximize carbon gain in understory environments and ability to regenerate
ruderal competition theory
high reproductive and growth rate on disturbed sites
stress tolerator competition theory
low reproductive and growth rate on undisturbed sites
competitor competition theory
low reproductive effort and high growth rate, productive sites