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Do individual organisms evolve during their lifetime?
No
Natural selection may affect individuals but only what evolves?
Populations
The product of inherited genotype and environmental influences
phenotype
What part of phenotype can natural selection act on?
Only variation with genetic component
Only natural selection causes organisms to adapt to an
environment
microevolution
a change in allele frequencies in a population over generations
sexual reproduction
can shuffle existing alleles into new combinations
gene duplications
duplicated genes can take on new functions by further mutations
germ cells
cells that mutations and duplications must occur in for natural selection to occur in
discrete characters
can be classified on an either or basis, this or that, no in between
quantitative characters
vary along a continuum within a population
discontinuous variation
distinct categories, tends to be qualitative, no in between
continuous variation
no distinct variation, no limit on the value, tends to be qualitative
height, weight, heart rate, finger length, leaf length
continuous variation examples
a lot of gene and environment between 2 extremes
continuous variation is controlled by
tongue rolling, finger prints, eye color, blood groups
discontinuous variation examples
a few genes limited by the number of phenotypes with no intermediates
discontinuous variation is controlled by
Genetic variation can be measured as
Nucleotide variability and average heterozygosity
nucleotide variability
is measured by comparing the DNA sequences of pairs of individuals
average heterozygosity
is gene variability, measured as the average percent of loci that are heterozygous in a population

What is the loci % that are heterozygous
4/18=22.2 percent, divide hetero by whole
geographic variation
differences between gene pools of separate populations
cline
variation that is a graded change in a trait alone a geographic axis
population
a localized group of individuals capable of interbreeding and producing fertile offspring of the same species
gene pool
consists of all the allele for all loci in a population
fixation
if all individuals in a population are homozygous for the same allele
p+q=1
the frequencies of all alleles
Hardy Weinburg principle
frequencies of all alleles and genotypes in a population remain constant from generation to generation
If a population violates the Hardy Weinburg principle
the population is evolving and is not in equilibrium from one generation to the next
p2+2pq+q2=1
genotype frequencies
The H-W equilibrium describes a hypothetical population that is
not evolving
The five H-W conditions are
no mutations
no natural selection
no sexual selection=must have random mating
no gene flow/no migration in and out of population
extremely large population size so there is no random affects
Are the conditons of hardy weinburg met in nature?
whats this called?
rarely met
evolution
Natural populations can evolve at some loci while
being in H-W at other loci
natural selection
differential success in reproduction results in certain alleles being passed to the next generation in greater proportions
the smaller the sample
the greater the chance of random effects cause deviation from predicted result
genetic drift
in small populations allele frequencies can fluctuate (drift) unpredictability from one generation to the next
Fixation in H-W
the establishment (fixing) of one allele in the population via the loss of other alleles leading to reduced genetic variation and increased homozygosity
bottleneck effect
a sudden reduction in population size due to a change in the environment
whats the result of the bottleneck effect?
the resulting gene pool may no longer be reflective of the original population’s gene pool
what happens if the bottleneck effect’s population remains small
it may be further affected by genetic drift
analogy for bottleneck effect:
Upside down bottle, big to thin
Humans can have bottleneck affect on species by
interferring with population size, number of alleles per locus
founder effect
when a few individuals become isolated from a larger population and seed a new population of heir own
allele frequencies in founder population can be different from those
in the parent population with fixation due to small population size
due to the founder effect, orginal will change frequences due to
population change and migration (violating hardy weinburg)
Effects of genetic drift
significant in small populations
causes allele frequencies to change at random
can lead to loss of genetic variation within populations\
can cause harmful alleles to become fixed
gene flow
movement of alleles among populations
gene flows via
emigration/immigration of fertile individuals or gametes
gene flow can increase or decrease the
fitness of a population depending on the environment and alleles introduced
natural selection is the only mechanism that
consistently causes adaptive evolution
new genetic variations arise by chance mutation (random) and these alleles
contribute to phenotypic variation
beneficial alleles are sorted and favored by natural selection in
enviornments
sorting
non-random
natural selection brings adaption to an environment by
acting on an organisms phenotype in that environment
are organisms perfect in their environment?
no, evolution is always occurring
reproductive success
different reproductive success via any difference that allows more offspring to be produced, mechanism of natural selection, depends on many factors
directional selection
favors individuals at one end or the other of the phenotypic range
disruptive selection
favors individuals at both extremes of the phenotypic range, average for both lines is the same.
stabilizing selection
favors intermediate variants and acts against extreme phenotypes
sexual selection
is the natural selection for mating success
sexual dimorphism
marked differences between the sexes in secondary sexual characteristics
male showiness due to a mate choice can
increase a male’s chances of attracting a female
intrasexual selection
competition among individuals of one sex, often males, for mates of the opposite sex
intersexual selection
often called mate choice, occurs when individuals of one sex, usually females, are choosy in selecting their mates
how do female preferences evolve? Hypothesis suggest
a trait that is related to male health, both the male trait and female preference for that trait should increase in frequency
how does female preferences evolve by chance?
female randomly develops a preference for the good genes male and in doing so both benefit
neutral variation
genetic variation that does not confer a selective advantage or disadvantage
Mechanisms that help to preserve genetic variation
diploidy
balancing selection
diploidy
maintains genetic variation in the form of hidden recessive alleles with heterozygotes
balancing selection
occurs when natural selection maintains stable frequencies of two or more phenotypic forms in a population
heterozygote advantage
frequency dependent selection
heterozygote advantage
when heterozygotes have a higher fitness than do homozygotes
frequency dependent selection
fitness of a phenotyoe declines if it becomes too common in the population
what happens when a frequency becomes too popular in a population/what’s favored?
the opposite
selection can only act on existing variations because
if no new beneficial mutations occur, then there are no new variations
evolution is limited by
historical constraints
adaptions are often
a compromise
not too good, not too bad
Chance and natural selection interacts with ___ and prevents perfect organisms
the environment