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abs. fitness (W) =
viability x fecundity
relative fitness (w) =
abs fitness of one / abs fitness of our reference genotype (biggest one)
selection coef (s) (selective disadvantage) (can be %)
1 - genotype relative fitness
relative fitness can ALSO =
1-s, 1+s

mean fitness of pop.

relative fitnesses
w11 w12 w22 (each abs fitness over biggest one)
new frequency of allele after selection

in positive selection

rate of adaptation
sp(1-p)
probability of fixation of a new beneficial mutation
2s
a beneficial mutation will increase in frequency from 10-90% in
4/s generations
s
strength of selection
q hat in overdominance
frequency of A2 at equilibrium

p hat in overdominance
equilibrium frequency of A1

mutation selection balance (dominant)
p hat = mu/s
mutation selection balance (recessive)
q hat = sqr root mu/s
equilibrium frequency in mutation selection balance
1- (mu/s)
fixation probability (new neutral)
1/2N
fixation probability (selected mutation)

|s| < 1/Ne
neutral
|s| > 1/Ne
removed by selection
|s| > 1/Ne LARGE
fixed by selection
phenotypic value =
genetic value + environmental value
breeders equation

narrow sense heritability

selection gradiet (beta)

change in mean phenotype (delta z hat)

phenotypic plasticity
genotype x environment interaction
number of nonsynonymous subsitution per nonsynonmous site
dN
under neutrality
dN = dS and dN/dS = pN/pS
dN/dS < 1
purifying selection
dN/dS > 1
positive selection
dN/dS > pN/pS
directional selection
dN/dS < pN/pS
purifying selection