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Atom
The smallest unit of an element that retains the element’s properties.
Molecule
Two or more atoms held together by chemical bonds.
Macromolecule
A very large molecule, such as a protein, nucleic acid, or polysaccharide.
Cell
The basic structural and functional unit of life.
Tissue
A group of similar cells that perform a common function.
Organ
A structure composed of two or more tissue types that performs a function.
Organism
An individual living entity.
Population
A group of individuals of the same species living in the same area.
Community
All populations of different species living and interacting in an area.
Ecosystem
A community plus its nonliving physical environment.
Biosphere
All regions of Earth where organisms live.
Evolution
Change in the characteristics of populations over generations.
Microevolution
Evolutionary change within populations, usually involving allele frequencies.
Macroevolution
Evolutionary change at or above the species level.
Species
A group of organisms that share a common gene pool and are reproductively compatible.
Molecular evolution
Evolutionary changes in DNA and protein sequences.
Empirical thought
Knowledge based on observations and experiments rather than speculation.
John Ray
Naturalist who developed an early biological species concept and classification system.
Carolus Linnaeus
Developed a hierarchical classification system and binomial nomenclature.
Georges Buffon
Early thinker who proposed that species could change and share common ancestors.
Jean-Baptiste Lamarck
Proposed evolution through use and disuse and inheritance of acquired characteristics.
Catastrophism
The idea that Earth's geological features resulted from sudden catastrophic events.
Uniformitarianism
The principle that the same gradual geological processes operate today as in the past.
Thomas Malthus
Argued that populations can increase faster than available resources.
Alfred Russel Wallace
Naturalist who independently developed the idea of evolution by natural selection.
Natural selection
Individuals with advantageous heritable traits leave more offspring.
Adaptation
A heritable trait that increases reproductive success in a particular environment.
Transitional forms
Fossils showing intermediate characteristics between ancestral and descendant groups.
Tiktaalik roseae
A transitional fossil with both fish and early tetrapod characteristics.
Pakicetus
Early whale relative showing the terrestrial ancestry of modern whales.
Odontoceti
Toothed whales, including dolphins and sperm whales.
Mysticeti
Baleen whales that filter-feed using baleen plates.
Biogeography
The study of the geographic distribution of organisms.
Endemic
Restricted to a particular geographic area.
Island fox
An endemic fox species found on the California Channel Islands.
Convergent evolution
The independent evolution of similar traits in distantly related organisms.
Analogous structures
Structures that have similar functions but evolved independently and do not share a common ancestral structure.
Selective breeding
The deliberate breeding of organisms with desired traits.
Artificial selection
Evolutionary change produced by humans selecting which organisms reproduce.
Homology
Similarity among traits resulting from common ancestry.
Homologous structures
Structures that share an underlying anatomical similarity because of common ancestry, even if their functions differ.
Vestigial structures
Reduced or modified structures inherited from ancestors that have little or no current function.
Developmental homology
Similarity in developmental patterns among related organisms resulting from common ancestry.
Molecular homology
Similarity in DNA or protein sequences resulting from common ancestry.
p53 protein sequence
A molecular sequence used as evidence of evolutionary relationships because similarities and differences in the protein reflect common ancestry.
Population genetics
The study of genetic variation and allele frequencies within populations.
Gene pool
All of the alleles present in a population.
Polymorphism
The presence of two or more genetic variants within a population.
Polymorphic gene
A gene that has two or more alleles in a population.
Monomorphic gene
A gene for which only one allele is present in a population.
SNP
A single-nucleotide polymorphism; a variation involving one nucleotide position in DNA.
Allele frequency
The proportion of a particular allele in a population's gene pool.
Genotype frequency
The proportion of individuals in a population with a particular genotype.
Homozygote
An individual with two identical alleles for a particular gene.
Heterozygote
An individual with two different alleles for a particular gene.
Hardy-Weinberg equilibrium
A condition in which allele and genotype frequencies remain constant from generation to generation in the absence of evolutionary forces.
Hardy-Weinberg equation
The equations p + q = 1 and p² + 2pq + q² = 1, used to calculate allele and genotype frequencies in a population at equilibrium.
p²
The expected frequency of homozygous individuals with two copies of allele p.
2pq
The expected frequency of heterozygous individuals with one p allele and one q allele.
q²
The expected frequency of homozygous individuals with two copies of allele q.
Hardy-Weinberg assumptions
Large population size, random mating, no mutation, no migration, and no natural selection.
Microevolution
A change in allele frequencies within a population over generations.
Natural selection
A process in which individuals with heritable traits that improve reproductive success leave more offspring.
Reproductive success
The relative contribution of an individual to the next generation through surviving and reproducing offspring.
Directional selection
Natural selection that favors individuals at one extreme of a range of phenotypes.
Stabilizing selection
Natural selection that favors intermediate phenotypes and selects against extreme phenotypes.
Diversifying selection
Natural selection that favors individuals at both extremes of a range of phenotypes.
Balancing selection
Natural selection that maintains two or more alleles in a population.
Heterozygote advantage
A form of balancing selection in which heterozygotes have greater reproductive success than either homozygote.
Negative frequency-dependent selection
Selection in which a phenotype or allele has greater fitness when it is rare.
Sexual selection
Selection resulting from differences among individuals in their ability to obtain mates or achieve reproductive success.
Intrasexual selection
Competition among members of the same sex for access to mates.
Intersexual selection
Mate choice in which individuals of one sex select mates based on particular traits.
Genetic drift
Evolutionary change in allele frequencies caused by random sampling of individuals.
Random genetic drift
Random changes in allele frequencies caused by chance events rather than differences in reproductive success.
Bottleneck effect
Genetic drift resulting from a drastic reduction in population size.
Founder effect
Genetic drift that occurs when a small group establishes a new population.
Neutral variation
Genetic variation that does not significantly affect an organism's reproductive success.
Gene flow
The movement of alleles between populations through the movement of individuals or gametes.
Random mating
A mating pattern in which individuals choose mates without regard to their phenotypes or genotypes.
Nonrandom mating
A mating pattern in which individuals choose mates based on particular traits or relationships.
Assortative mating
Nonrandom mating in which individuals with similar traits are more likely to mate.
Disassortative mating
Nonrandom mating in which individuals with different traits are more likely to mate.
Inbreeding
Mating between individuals that are more closely related than randomly selected individuals in a population.
Inbreeding depression
Reduced reproductive success resulting from increased homozygosity caused by inbreeding.
Speciation
The evolutionary process by which new species arise.
Morphology
The form and structural characteristics of an organism.
Reproductive isolation
The inability of populations to successfully interbreed and produce fertile offspring.
Species concept
A definition or framework used to identify and distinguish species.
Biological species concept
The concept that defines a species as a group of actually or potentially interbreeding natural populations that are reproductively isolated from other such groups.
Evolutionary lineage concept
The concept that defines a species as a lineage of populations that maintains its identity from other lineages and has its own evolutionary tendencies and historical fate.
Lineage
A sequence of populations or organisms connected through ancestor-descendant relationships.
Ecological species concept
The concept that defines a species according to its ecological niche.
General lineage concept
The concept that defines a species as a separately evolving lineage of organisms or populations.
Reproductive isolating mechanisms
Biological mechanisms that prevent populations from successfully reproducing with one another.
Interspecies hybrid
An offspring produced by reproduction between individuals from two different species.
Habitat isolation
Reproductive isolation that occurs when populations live in different habitats and therefore rarely encounter each other.
Temporal isolation
Reproductive isolation caused by populations reproducing at different times.
Behavioral isolation
Reproductive isolation caused by differences in courtship or other reproductive behaviors.
Mechanical isolation
Reproductive isolation caused by differences in reproductive structures that prevent mating or pollen transfer.