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Behave
A response to stimuli from enviroment
contruibutes to homeostasis
Essential to fitness of a species
Individual contribution to gene pool
Essential to aquiring nutrients; use of recources
Essential to mating/ maintaning a speices
Innate Behavior
Born with it (genetic component)
Requires no learning
Nearly all in a speices exhibit it
Learned Behavior
Modifieed by experience
Requires diff. stimuli
Causation
Is the “how” and “why” of behavior
Proximate causation
The “how” - Physiological component ( sex hormones adrenaline “fight or flight”) can include nature and nurture.
Ultimate causation
the “why”- evolutionary component. Natural selection at work (mating rituals, survival behavior).
Genetic Component
Maze savvy rats mated with maze savvy rats yeild maze un-savvy rats ( opposite ture as well)
Genetically identical twins have identical genome- separated at birth then reunited later in life: similar temoerment personalities, interests.
In our lab- c. elegans & the genes for learning.
Mice have gene responsible for maternal care. was isolated & disabled in lab: mice no longer cuddled & kept young warm.
Enviorment component (“nurture”)
cross-fostering studies: one species placed in care of another (much different) at very early age- as get older tent to display some of other species characteristics.
Fixed action patterns
sequences of unlearned acts directly linked to a simple stimulus called a sign stimulus (ex. geese egg rolling).
Migration
Long-range 2-way movements,
celestial navagation (sun/ North Star)
when day/ night may use circadian clock
if cloudy may use earth’s magnetic field (homing pigeon/ magnet mangnetite in head of migrating species
Communication
Recognizable visual, auditory or chemical stimuli that elicit a response.
Speices recognition
courtship displays recognized by 1 species
blue footed booby
albatross
helps prevent mating between species/ isolate genes with in a speices
actions initiate stimulus- response chain
Dancers & dialogue (honeybee dance laugage calling to warn of danger
Pheromones
Chemicals used to communicate location, danger, sex attractants during mating season
Learned Behavior
Behavior modified by experiences
2 types: non-associative & associative
Non-associative learning
Habituation
(simplest type of learning)
ex. work at rocky quarry with frequent explosions jump at first eventually no reaction
Associative learning
Connection between a stimulus and a response
Imprinting
a mix between innate and learned behavior
Learned social interactions between different species
Has a critical period where attachment forms
Classical conditioning
Ivan pavlov & pavlov’s dogs
a paired association between 2 diff. types of stimuli
the animal forms association
the experiment:
meat powder presented to dog, dog salvates (unconditioned stimulus= meat powder, unconditions respond= salivates)
bell rung at time of presenting meat
eventually only the bell ringing caused dog salvation
Operant Conditioning
“Trial and error” learning B.F. Skinner & the “Skinner Box”
animals learn to associate a reward with behavior
very important learning for most vertabretes and in the “wild”- natural enviroment for feeding (training fog, bird, catching butterfly
Spatial Learning
Learning to use landmarks to help find food , nests, ect.
Using the landmarks, a cognitive map is created
prevents need from moving from landmark to landmark
ex. Ground bee nest
Honey bee waggle dance (a combination of behaviors)
Associations that animals make likely to be ones they come across and provide selective advantage
Pill Bugs
Terestrial crustations (breathe with gills)
decomposers
example organisms for orientation movement
Taxis
Movement directly toward or away from a stimulus
positive taxis= towards
Negative taxis= away
Kinesis
Random movement in an enviorment- undirected “searching” for desired stimuli. If kenisis results in disired or undesired stimuli can become taxis.
Cognition and Problem solving
The most complex types of learning requires:
Awareness
Reasoning
Recollection
Judgement
This type of learning can be social
Juvinials observing the experienced
can involve a sensitive period where the learning is critical to take place
if miss critical period wont learn properly
Foraging behavior
Not only finding food but energy expended finding it (distance to, battle to capture/ kill, risk of injury/ death in process)
Optimal foraging model
Natual selection favors behaviors minimizing cost & maximizing benefit.
frruit fly gene analysis (forager gene) provides this low-density “siter” allele high density “rover” allele.
Mating behavior
More than a partner; also carring for offspring, competition for mates, most genetically “fit” partner
most species promiscuous- no pair bonding, many partners
ensures offspring/ survival of species; care of young may be compromise
if pair bonding occurs can be either:
monogamous
polygamous
polygyny
polyandry
Monogamous
One male & one female don’t generally display sexual dimorphism (size/ color/ ornamentation diff.)
Polygamous
1 individual of one sex mating with several of other sex
Polygyny
most common- one male many females
Polygandry
less common- one female many males. Morphologic diff. seen in females
Parental care
Young helpless & need constant food- often monogamous relationship (one guards other hunts)
Polygamy
Gets down to “certainty of paternity” or “whos your daddy”
basically if you know they’re your offspring your more likely to care for them
Aitruism
Species can exhibit “selflessness”
reduce an individuals fitness (passing his/her genes on to the next generation) by putting oneself in danger.
increases fitness of other individuals in population
ex. ground squirrle sees hawk stays above ground to warn others- risk of predation but others safe
Inclusive fitness
total effect individual has on sending its genes on to the next generation
Visual signals
color & mating behavior/ color and danger
Audible signals
warning call, mating song
Tactile signals
touch- part of mating rituals
Electrical signals
electric fishes- sharks/ rays others can generate weak electrical signals
Chemical signals
Pheromones- for locomotion/ mating/ teritory
Community
Assemblage of populations living close enough for possible interactions
Individualistic concept
nothing more than species living in the same space at the same time
some recent research supports ( with global warming)
Holistic concept
many accept currently- speicies function as an integrated unit. Need all to function properly
Community Interactions
Interactions with in a community are interspecific.
can do 3 things:
Help those involved (+ effect)
Harm those involved (- effect)
Have no effect on those invoulved (0 effect)
Commpetition
((-/-) both harmed in some way) species can coexist but commpetition for recourses will limit growth, survival, & space they occupy. (weeds, lawn)
Compettitive exclusion
if 2 species compete for the same limiting recource, both can NOT coexist
Niche
All the ways an organism uses it’s recources (biotic & abiotic)
Fundemental = what could occupy in perfect world
realized=what it does occupy
Species can not coexist in a community of their niches are identical
Recource partitioning
enambles similar species to occupy same area using different set of recources.
predation
(+/-) predator kill & eats prey. Predation skills and prey avoidance skills refined through natural selection- need reproductive success.
Predator adaptations
acute senses, claws, teeth, poison, agility
Prey adaptations
cryptic coloration- camoflouge: aposematic coloration- bright colors associated with toxins produced or ingested & expressed by prey.
Mimmicry
two resemble each other
batesian= harmless mimics harmful
mullarian= 1 harmful mimics markings of another harmful
Herbivory
(+/-)
An herbivore eats plants of alge
has led to evolution of plant mechanical (spines/ thorns) & chemical (toxins) defenses & adaptations by herbivores
Symbiosis
2 or more speices living in initmate contact (very close/ direct)
Parastitism
(+/-) 1 organism(parasite) derives nourishment from another organism (host) which is harmed in the process (ticks, tapeworms)
Mutualism
(+/+) interaction that benefits both species (lichen, mycorrhizae, termites & digestion)
Commensalism
(+/ 0) 1 species benefits & other is neither helped nor harmed - hitch hiker barnacles
Facilitation
(+/+) or (+/ 0) 1 species cna have positive effects on another species with out direct intimate contact (black rush in salt marshes)
Ecological succession
Trend in species & diversity of an area from simple to complex. Different speices alter the enviorment, allowing others to take hold.
Primary succession
occurs on bare, lifeless substrate ( rock, sandy beach, volcanic lava bed)
Pioneering species
Are first speices ot inhabit area
must be very tolerant of harsh abiotic conditions
Secondary Succession
occurs in areas that have been disturbed - soil remainns intact (fire, clear- cutting for logging/ forestry)
Tolerance
(1st to inhabit) pioneering organisms must be hardy / able to withstand abiotic conditions
Facilitation (part of ecological succession)
Pioneering (or early species in case of secondary Succession) alter abiotic conditions (soil/ PH) facilitation growth of less tolerant organisms
Inhabition
1 speices that facilitated the growth of another may eventually be out- competed (growth inhabited) by another speices
Speicies Diversity
Speices diversity of a community is the variety of organisms that make up the community.
2 components: Speices richness & relative abundance
Speices richness
is the total number of different species in a community
Relative abundance
is the proportion each species represents of the total individuals in the community
Ecotone
2 ecosystems overlap.
communities with a higher diversity are:
more productive & more stable in their productivity
better able to withstand & recover from enviromental stresses
more resistant to invasive species (not native) organisms that become established outside their native range
Trophic levels
shows feeding relationships between individuals with in a community
Huge cey factor in community dynamics
Decomposers “feed” on every trophic level
Food web
Is a branching food chain with complex trophic interactions
Energy Hypothesis
suggests that length is limited by inefficient energy transfer (about 90% lost at each level, supports 10%)
Dynamic stability hypothesis
long food chains are less stable than short ones.
fluctuation at lower trophic levels are magnified at upper ones
Dominant Species
the most abundant or the most biomass in a community
may be able to outcompete for limited recources
may have afaptioms to better avoid disease / predation
Keystone species
Have profound effects on communities
some construct their own ecosystems (beavers)
some profound effects numbers at each trophic level
Bottom- up model
Proposed an undriectional influence from lower to higher trophic levels
presence or absence of mineral nutrients determins primary producers which influence all other trophic levels
Top- down model
(AKA trophic Cascade) proposes that control comes from the trophic level alone
predators control herbivores which in turn controls primary producers
Biomani Pulation
Human influence on speices at different trophic levels to help nestone polluted communities
Primary productivity
The rate at which photosynthetic organisms build organic compounds (fix atmospheric carbon into sugars) & synthesize oxygen.
Gross primary productivity (GPP)
total amount produced in any given period
Net primary productivity (NPP)
total amount produced minus what plants use for respiration
* Yes plants have a mitochondria & do cell respiration @ hight, low sun, and during points of hight ATP demand
Population Ecology
A population is a group of individuals of a single species living in the same general area
populations are described by their boundaries & size
Density
Is the number of individuals per unit or voulume (if in liquid- bacteria or algae)
Dispersion
is the pattern of spacing among individuals withing the boundaries of the population
Density is result of interplay between processes that add individuals to a population & those that remove individuals
Immigration
is the influs of new individuals dorm other areas
Emigration
is the movement of individuals ou of population
Population size
In most cases, impractical or impossible to count all individuals in population
sampling techniques used to estimate densities & total population sizes
Mark- recapture method
scientice capture, tag & release a random sample of individuals (s) in a population
marked individuals are given time to mix back into the population
Scientists capture a second sample of individuals (n) & note how many of them are marked (x)
population size (N) is estimated by: N=(sn/x)
Clumped (dispersion)
Individuals aggregate in patches
common clumped dispersion may be influenced by recource availibity (uneven or scarce) and social behavior (travling together)
Uniform (dispersion)
evenlu distributed
may be influenced by social interactions such as territoriality the defense of a bounded space against other individuals (compettition for recourses)
common when closely spaced plants compete for recources (water, sunlight)
some plants (walnut) secrete toxin into soil to prevent others
Random (dispersion)
the position of each individual is independent of other individuals
Not common unless verry little species interaction
Species survival
can be influenced by biotic or abiotic factors
Abiotic are “non-living” interactions- many times enviormental interaction
Biotic are living interactions
Density Dependent
Result of more individuals in population is an example of negative feedback control over population.
population increases
mortality can increase
reproduction rates can deline
Density Independent
Factors unrelated to population that affected growth
often enviormental factors like drought, cold winter, hurricane, volcano.
cause growth curve of population to appear erratic not a smooth curve
Exponential Growth Model
An idealized situation with abundant recources & reproduction at Physiological capacity
allows for understanding about a species potential capacity to increase & the conditions required for it
immigration= emmigration
R-selected speices
Births exceed deaths
J shaped curve
Logistic Growth Model
Tru J-shaped curve unrealisitc in most cases
immigration not equal emmigration
Not infinite recources
Population rate will decrease as pop. reaches carying compacity (denoted with a “k” on graphs)
Predator- prey interactions
many times speices populations interconnected
snowshoe hare feeds on willowbark. if quantity decreases hare feeds on high fiber, birth rate reducing diet. this follows 10 year cycle (rise/ fall in pop. rates)