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Species richness
The number of different species present in a community; simple count, does not consider abundance differences (not number of individuals)
Equitability (evenness)
How individuals are distributed among the species present
Low
If one species makes up 90% of individuals, evenness is ____
High
If all species have roughly the same number, evenness is ____
High diversity
Many species; grow in roughly equal abundance
Shannon index
Rages from 0 to infinity
Simpsons D index
Ranges from 0 to 1
Shannon index
H’= -sum of (pilnpi)
Pi
Shannon index: Proportion of individuals in x species
~0.5
Shannon index: low diversity
~1.5-2.5
Shannon index: Moderate diversity
>2.5
Shannon index: High diversity
Simpsons D index
D=sum of (pi2)
Low diversity
Simpsons D index: If one species dominates (90%) D will be close to 1 (___ ___)
High values
Simpsons D index: Low diversity
Low values
Simpsons D index: High diversity
Habitat diversity
More types of habitat, more niches (mountains have different zones of temps and moisture creating microhabitats and niches)
Tropical regions
Over half of all species occurs in ~1% of Earth’s land
Time; area; large; less; larger
Tropical diversity (___ and ___): long-term stability and ____ continuous areas promote speciation; Great tropical areas allows large species ranges and ___ extinction; ___ areas= more individuals, and greater genetic variation
Conservatism
Tropical diversity (____): Many species originated in the tropics, and never left; adaptations to warm, wet climates makes it hard to colonize temperate regions; Most lineages still live near their evolutionary origins
Productivity
Tropical diversity (____): More sunlight and more rainfall in tropical areas allow for more plant biomass; increased photosynthesis; high primary productivity supports more complex food webs; abundant food means less extinction risk
Spatial heterogeneity
Tropical diversity (___): Variation in habitat (light, temperatures, moisture) provides many niches; variation prevents one species from dominating
Evolutionary time
Tropical diversity (___ ___): tropics are extremely old; allows for more time to evolve and diversify
Evolutionary speed
Tropical diversity (___ ___): Evolution occurs faster in warm climates; mutation increases with temperature; faster reproduction=more generations per time unit (faster evolutionary change)
Climate stability
Tropical diversity (___ ___): Tropical climates vary less then temperate climates; decreases extinction and allows niche specialization
Disturbance timing
Tropical diversity (___ ___): How often disturbance happens
Too little
Disturbance timing: Competitive exclusion (one species dominates)
Too much
Disturbance timing: Species cannot establish
Intermediate
Disturbance timing: Mix of colonizers (pioneers) and specialists coexisting
Weather
Current state of the atmosphere at any given time
Climate
Long-term weather conditions; includes variability
Climate change
Directional change in climate over a period of time
3 decades
Climate change has a directional change over a period of time of at least # ____
1.1
Average global surface temperature increased by ___ C between 1880 and 2020
Glacier; sea; extreme; shift; insecurity
Current climate change: ___ loss; ___ levels rise; ___ weather; ecosystem ___; food and water ____
Dryer
Climate change: Deserts are getting ___
Wetter
Climate change: Temperate areas are getting ____
Anthropogenic
Cause of climate change through fossil fuels, deforestation, and agricultural processes (humans)
Fossil fuel burning
Coal, oil, natural gas
Agricultural processes
Changes the amount of sunlight that is absorbed/reradiated back into space; greenhouse gasses (trap heat on Earth)
Food chain impacts
Impacts of climate change: Polar bears hunt seals, but now seals surface in open ocean instead of holes in ice
Ocean acidification
Impacts of climate change: Increases carbon dioxide levels; coral bleaching
Terrestrial organisms
6% of insects, 8% of plants and 4% of vertebrates will loose at least ½ of their geographic range with 1.5C increase
Range shifts
Species have to move to more tolerable climates
Shifts threaten species
Cannot move fast enough; have nowhere to move; paths are blocked by human structures
Surface; water vapor; growing; sea; temperature
Climate change: increased ____ temperature; increased __ __ in atmosphere; increased ___ season; increased ___ level; increased sea ____
Snow; ice sheets; sea
Climate change: Decreased ___ cover; decreased ___ ___ and glaciers; decreased ___ ice
1.5; fossil fuels
Current goal: limit warming to ___ C; reduce use of ___ ___
Extinction
Permanent loss of a species
Mass extinctions
Needs timing, multiple ecosystems, and numbers to classify as this
5
Earth has had __ mass extinctions
Natural
Extinction is a ___ part of Earth’s history
Scale; time
Extinction becomes concerning when ___ increases and ___ decreases
10-100
Normal rate (extinction): roughly __-__ species per year
Background extinction
Normal, ongoing loss of species over geological time
Mass extinction
Widespread, rapid loss of biodiversity; numerous groups and numerous ecosystems
Fossil records
Usually across strata in rocks (geological boundaries)
Geochemical; volcano; impact
Identification of extinction “cause”: Changes in ____ makeup; presence of ___ deposits, ___ sites
Large scale volcanism
Mass extinctions: Release greenhouse gases and other compounds; rapid warming or cooling, acid rain, ocean acidification
Asteroid impacts
Mass extinctions: Shockwaves, fire, dust, aerosols, rapid cooling, food-web collapse
Climate change
Mass extinctions: Alters habitats faster than many species can adapt
Ocean anoxia
Mass extinctions: Decreased oxygen content
Niche vacancies
After extinction: Survivors may occupy habitats left empty
Adaptive radiation
After extinction: Surviving lineages diversify into multiple new species
Recovery period
After extinction: Long term recovery to approach previous levels of diversity and primary productivity; changes in abundance and diversity
~444mya
5 mass extinctions: End-Ordovician
~359-372mya
5 mass extinctions: Late Devonian
~252mya
5 mass extinctions: End-Permian
~201mya
5 mass extinctions: End-Triassic
~65mya
5 mass extinctions: End-Cretaceous
End Ordovician
Loss of majority of marine organisms; formation of massive ice sheets; sea levels dropped; followed by rapid warming
Global cooling
End Ordovician mass extinction happened due to rapid ___ ___
Double pulse extinction
The rapid global warming following the end Ordovician mass extinction caused a ___ ___ ___
Late Devonian
~75% species loss; marine life; armored fishes; expansion of land plants increased runoff
Ocean oxygen; plants; volcanism
Late Devonian mass extinction caused due to reduction in ___ ___; expansion of land ___ (increased chemical weathering; increased nutrient in waterways; increased algae content decreased carbon dioxide and oxygen levels); potential ____
End Permian
Lasted 50,000 years; the great dying; ~96% marine, 70% terrestrial; 50% of families went extinct
Lava flows; atmosphere; ozone; warming; acid; acidification; hurricanes
End Permian mass extinction caused by huge __ __ from Russia (released massive amounts of CO2); toxic ___ (from CO2 and volcanic gas); loss of ___ layer; global ___; ___ rain; ocean ___; Mega ____
End Triassic
~75%; large amphibians, marine reptiles
Pangea; warming
End Triassic mass extinction caused by CO2 released from ____ splitting; global ___
End Cretaceous
~75%; non-avian dinosaurs; plesiosaurs; marine plankton; marine reptiles; non-seed plants; Survivors: Deep sea organisms
Outcompete; glaciation; volcanic
End Cretaceous mass extinction theories: Mammals ___ dinosaurs; ____; ____ activity
Asteroid impact
End Cretaceous mass extinction caused by ___ ___
Before; diversify; marine; partial
Mammals outcompete: Mammals existed ____ extinctions; mammals ____ after dinosaurs went extinct; Doesn’t explain ___ extinctions (___ explanation, but not primary cause)
Mobile; globally
Glaciation/climate cooling: BUT dinosaurs are ___ and could leave cooling areas; Extinction affected organisms ___, not just in areas that cooled
CO2; acid; decline; contributed
Volcanic eruptions: increased ___; lead to ___ rain; stress systems and cause ecosystem ___; BUT too ___ to explain sudden extinction peak (___, but not sole cause)
Seeded; crater; large
Asteroid impact: ___ plants survived (can go dormant in the dark); ___ matches size and age of event; Global extinction pattern: must have been something ___
Time 0
After asteroid: Instant firestorms; Ejected materials re-entering atmosphere; mega-tsunamis over km tall near crash site; massive earthquakes
Minutes/days later
After asteroid: Dust/soot/ aerosols block sunlight; photosynthesis halts; rapid global cooling
Weeks/years later
After asteroid: Food web collapse; plankton crash led to marine die-off; herbivores starve, carnivores crash; fungi and detritivores thrive; ferns=pioneer species
Competition; predation; disease; environmental change; cataclysms
Mass extinctions aren’t always dramatic; ____ (invasive species; early mammals); ___ (cats); ___ (brought by humans); ___ ___ (long term shifts); ____ (asteroids, volcanic eruptions, floods, ocean anoxia)
Dutch elm disease
Extinction event (disease): fungal pathogen; spread by beetles; destroyed native elms in Europe and north America
Terminal Pleistocene extinction
Is a major extinction even NOT a mass extinction
Terminal Pleistocene extinction
~65-70% of large animals in north america; ~50,000-10,000 ya; primarily affected larger megafauna (>100lb) [mammoth, mastodon, sabretooth, giant ground sloth, giant birds, horse, camels, dire wolves]
Human hunting; climate change; recent
Terminal Pleistocene extinction caused by ___ ___ pressure (new hunting tools and techniques; lack of fear in organisms); ____ ___ led to rapid warming; Most ___ large-scale extinction