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Water will always move _______ concentration gradients
magnitude gradients
Determines if organism loses or gains H2O from environment
evaporation
Accounts for much of water lost by terrestrial organisms
Relative humidity equation
Relative humidity= (water vapor density/saturation water vapor density)*100
Water vapor density measurement
measured as the water vapor per unit volume of air
saturation water vapor density measurement
measured as quantity of water vapor air can potentially hold, changes with temperature
Total atmospheric pressure
pressure exerted by all gases in the air
Water vapor pressure
partial pressure due to water vapor
Saturation water vapor pressure
pressure exerted by water vapor in air saturated by water
Vapor pressure deficit
difference between WVP and SWVP at a particular temperature
Higher vapor pressure deficit causes ____________ rate of evaporation
higher
Is water more concentrated in freshwater environments or in the oceans?
freshwater environments
osmosis
diffusion across a semipermeable membrane
Isosmotic (isoosmotic)
body fluids in organism and external fluid are at the same concentration
Hypoosmotic
body fluids are at higher water concentration than external environment
Hyperosmotic
body fluids are at a lower water concentration than external environment
Water potential (psi)
capacity of water to perform work, generally negative in nature, water potential of the solute measures the reduction in water potential due to dissolved substances
0
water potential of pure water
Matric forces
waters tendency to adhere to container walls
Ψpressure
reduction in water potential due to negative pressure created by water evaporating from leaves
2 major challenges terrestrial organisms face
-evaporative loss to environment
-limited access to replacement water
Metabolic water
the water released during cellular respiration
What will a plant allocate resources to?
the structure or function responsible to the acquisition of resources that are most limiting
water conservation by plants and animals
•Many terrestrial organisms equipped with waterproof outer covering.
•Concentrated urine / feces.
•Behavioral adaptations to avoid stress times.
•Drop leaves in response to drought.
•Thick leaves
•Few stomata
Periodic dormancy
Autotrophs
use inorganic source of C and energy
Photosynthetic autotrophs
use CO2 as C source and sunlight as energy
Chemosynthetic autotrophs
use inorganic molecules as source of C and energy
photon flux density
number of photons striking surface area
What factors influence the distribution of light quality/quantity across the earths surface?
*water depth
*water quality
*elevation
*shade
*atmosphere quality
*aspect
*burrows and rocks
Albedo
reflectivity of surfaces (such as soil)
Where do plants get their carbon?
from the atmosphere
How does carbon enter the plant?
CO2 enters the stomata of plants
C3 photosynthesis
named after PGA (3 carbon acid-3-phosphoglycerate), used by most plants and algae
Rubisco
the most abundant enzyme on earth, needs lots of CO2, the enzyme that catalyzes CO2 fixation (low affinity for CO2), catalyzes photorespiration
Photorespiration
O2 + RuBP
occurs in the light
consumes energy
C4 photosynthesis
4 carbon acid produced, PEP carboxylase (enzyme), acids produced during carbon fixation go to specialized cells called bundle sheath, there C3 pathway ensues
CAM photosynthesis
limited to succulent plants in arid and semi-arid environments, carbon fixation takes place at night, reduced water loss, low rates of photosynthesis, extremely high rates of water use efficiency
Denitrifiers
convert nitrate to dinitrogen gas (which can be used by organisms), need NO3
herbivores
mostly feed on plants
carnivores
mostly feed on animal flesh
detritivores
mostly feed on non-living organic matter
What 5 elements make up 93-97% of biomass of plants, animals, fungi, and bacteria
C, O, H, N, P
N and P are usually the hardest to get
How is food quality measured
C:N ratios
Plant defense mechanisms
*thorns/spines
*grow tall
*enlist the help of other organisms for protection
*cellulose
*lignin
*silica
*toxins
*digestion reducing compounds
Organisms prefer eating things with _____ C:N ratios
lower
What constitutes most of the biomass on earth?
plants
How do animal prey not get eaten?
*poison
*venom
*camouflage
*hard outer shells
*detaching limbs
*run fast
*climb well
*nocturnal
*spines
*stay in groups
*emitting odor
*Aposematic coloring
*Mullerian mimicry
*Batesian mimicry
Aposematic coloring
warning colors
Mullerian mimicry
co-mimicry among several spp. of noxious organisms
Batesian mimicry
harmless spp. mimic noxious spp.
Predators
selection agents for refined prey defense
Evolutionary arms race
predator and prey race to evolve to determine the fate of the ecology of the species
Rate of photosynthesis increase with ________
photon flux density
Photosynthetic response curves
increase and then level off at a maximum photosynthesis rate
3 basic functional responses
-feeding rate increases linearly as food density increases then levels off (consumers require little or no search and no handling time)
-feeding rate rises in proportion to food density (feeding rate partially limited by search/handling time), most common
-slow at first then rate increases most rapidly at intermediate densities (s-shaped) (organism learns how to eat diet item more efficiently)
Optimal foraging theory
organisms often have options/choices regarding what they feed on, prey selection maximizes energy intake rate
optimal foraging theory equation
E/T=Ne1E1-Cs/1+ (Ne1H1)
E=Energy
T=Time
Ne1=Number of diet item 1
Cs=Cost of searching H1
H1=Handling time
Behavioral Ecology
Interactions btwn. organisms and environment mediated by behavior
Sociobiology
examines social behavior especially as it relates to how behavior relates to fitness
Female
produces larger, more energetically costly gametes
Males
Produce smaller, less energetically costly gametes
Resource access
what limits reproduction
mate access
what limits male reproduction
sexual selection
differences in reproductive rates in individuals as a result of differences in mating success
Intrasexual Selection
individuals of one sex compete among themselves for mates (ex. elk fighting)
Intersexual selection
individuals of one sex choose mates among members of opposite sex (ex. lyre bird dancing)
genetic variation
what evolution acts on
sociality
degree to which individuals in a population associate in social groups
*cooperative feeding
*defense of the social group
*restricted reproductive opportunities
evolution of sociality is generally accompanied by:
eusociality
more complex level of sociality
*individuals of >1 generation living together
*cooperative care of young
*division of individuals into non-reproductive and reproductive castes
characteristics of eusociality
*inclusive fitness
*inherited territory
*kin selection
benefits of cooperative breeding
inclusive fitness
improve survival and reproductive rates of family members
kin selection
evolutionary strategy that favors the reproductive success of an organisms relatives even at cost to its own survival and reproduction
Hamilton’s rule
selection will favor diverting resources to kin when benefits to the helper exceed costs
Hamilton’s rule equation
RgB - C > 0
Rg=degree of relatedness
B=reproductive benefit gained by the recipient of the help.
C=costs
population
group of individuals of a single species inhabiting a specific area, characterized by the # of individuals and their density
number of individuals, density, age distributions, growth rates, distribution, and abundance
characteristics of a population
niche
summary of factors that influence growth, survival, and reproduction of a species
niche- hutchinsons definition
n-dimensional hyper-volume
n equates to # of environmental factors important to survival and reproduction of a species. volume=range
fundamental niche
hypervolume (abiotic factors)
realized niche
biotic interactions such as competition or symbioses that may restrict or expand environments where species may live
food production; water supply; habitat; incidence of parasites, pathogens, and competitors; human impacts
climate often influences species distributions via
random distribution
equal chance of being anywhere, uniform distribution of resources
regular distribution
uniformly spaced, exclusive use of areas, individuals avoid one another
clumped distribution
unequal chance of being anywhere, mutual attraction between individuals, patchy resource distribution
rarity I
extensive range, broad habitat tolerance, small local populations (ex. peregrine falcon)
rarity II
extensive range, large populations, narrow habitat tolerance (ex. passenger pigeon)
rarity III
restricted range, narrow habitat tolerance, small populations (California condor)
metapopulations
groups of subpopulations living on patches of habitat connected by an exchange of individuals
cohort life table, static life table, age distribution
3 main methods of estimation
cohort life table
identify individuals born at same time and keep records from birth (best, but difficult)
static life table
record age-at-death of individuals
age distribution
calculate difference in proportion of individuals in each age class, assumes difference from mortality
Type I survivorship curve
majority of mortality occurs among older individuals (ex. dall sheep)
Type II survivorship curve
constant rate of survival throughout lifetime (ex. american robins and mud turtles)
Type III survivorship curves
high mortality among young, followed by high survivorship (ex. perennial shrub)
anticipate future populations, identify the drivers of population growth, manage populations of conservation concern, evaluate interactions among species
modeling population growth used to:
generations do not overlap
growth can be modeled geometrically when
Geometric growth equation
Nt = Noλt
Nt = Number of individuals at time t.
No = Initial number of individuals.
λ = Geometric rate of increase. (# offspring left by individual in 1 time interval)
t = Number of time intervals or generations.