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Fossil Record
evidence of life preserved in a geologic context
body fossils, trace fossils (Ichnofossils)
Fossils provide information about:
anatomy
feeding strategies
geographical distribution
ecological relationships
past climates and environments
Type of Fossils: Body Fossils
examples includes:
bones and teeth
shells and exoskeletons
wood
remains can represent a whole organism or just parts of an organism
remains may be in one piece or granted/scattered
more rarely:
skin, soft tissue, and organs
leaves and flowers
Types of preservation:
Typical:
unaltered skeleton
per mineralization, pores become filled with silica
replacement with another material like quarts or pyrite
molds and casts. original material is dissolved and sediment fills the space
carbonization, preserving as carbon films
Exceptional:
fossil lagerstätten:
konzentrat (concentration) lagerstätten: anomalously high amounts of fossil material
konservat (conservation) lagerstätten: unusually well-preserved fossils, including soft parts
stagnation deposits (anoxia, low current flow)
obrustion deposits (very rapid burial)
conservation traps (amber, for example)
Types of Fossils: Trace Fossils (Ichnofossils)
we don’t necessarily know the identity of a trace maker
taxonomy: Ichnogenera and Ichnospecies
provide information about:
how an organism moved
feeding patterns
burrowing patterns in sediment
community structure
examples:
footprints, trackways, and resting traces
burrows and tunnels
feeding traces
coprolites
Fodinichnia- Feeding burrows
branching burrow traces of a deposit-feeding invertebrate
e.g. Chondrites
precambrian- Recent
environmental settings: fully marine conditions perhaps indicative of low oxygen zones
Fodinchnia- feeding burrows
burrow traces of a deposit feeing invertebrate
e.g. Teichichnus
cambrian- recent
environmental settings: found in lower marine shore faces, intertidal settings, lagoons, deep-marine fans and abyssal plains, and brackish water settings like marine estuaries
Domichnia- Dwelling burrows
unbranched burrow traces of a suspension-feeding invertebrate
e.g. Skolithos
precambrian- recent
environmental settings: found in virtually every type of environment from marine to terrestrial present in high-energy marine conditions, especially in nearshore shallow Marien facies, deep marine submarine channels and canyons; continental floodplain and dune settings
Domichnia- Dwelling burrows:
straight or gently curved burrows, horizontal to oblique bedding planes
e.g. Planolites
precambrian- recent
environmental settings: shallow marine- deep marine deposits to continental. found in alluvial, lacustrine, and eolian settings
not only animals produce trace fossils
rhizolithi (plant root structure)
analysis of track traces
different parts of a track can create different traces
type of substrate, and texture of sediment is important
can reveal details about the track maker’s anatomy and movement
dinosaur footprints and trackways
gives clues about locomotion styles
gives clues about dinosaur families and communities
give clues about predation behaviors, for example maybe some dinosaurs hunted in groups

One organism can produce many traces based on their behavior
a trace fossil may look different depending on how it was produced (for example the interaction between trace maker and sediment) and how it is viewed
traces that resulted from animal activity
e.g. deer, hog, horse
Repichnia- Locomotive traces
e.g. Chirotherium from the Triassic
pentadactyl footprints
not obvious who the producer was
Adolf Seilacher
Seilacher’s interest included trace fossils, evolution of invertebrates and exceptional preservation (Lagerstätten)
Stuttgart

Tubingen

Changes in setting reflected in changes in observed traces
Seilacher created a naming system based on using ichnotaxa to describe trace fossil types
Cubichnia= Resting traces
Domichnia= Dwelling traces
Fodinichnia= Feeding traces
Pascichnia= Grazing traces
Repichnia= Locomotory traces
lots of other terms have since been created
Seilacher also developed the frameworks of constructional morphology and morphodynamics

“Taphonomy” coined by Ivan Efremov in 1940
= “The study of the laws governing the transition (in all its details) of organic remains from the biosphere into the lithosphere”
also includes discovery, collection bias, and curation of remains
natural sampling of biotas
occurrence in the fossil record also depends on species diversity, morphological disparity, and Ease of preservation
differing preservation potentials
A collection of filters
the study of the transition from the biosphere to the lithosphere
sources of bias:
biochemical
anatomical
ecological
geological
Sequential stages of taphonomy:

Assemblages:
Biocenosis (life assemblage)
organisms have the correct orientation
trace fossils may be present
different organisms from the same community are present
mature and juvenile individuals may be present
Thanatocenosis (death assemblage)
organisms are of similar size and/or shape
organisms may be from different ecological communities
Autochthonous (In situ)
Allochthonous (Transported)
Single-event (census/catastrophic) assemblages:
unusual events like sudden anoxia, severe storms, droughts, and volcanic eruptions that kill large numbers of individuals at one time. short term, snapshot
Attritional (time-averaged) assemblages:
discarded organic products (like pollen and leaves and input from normal mortality over time. Long term
Precambrian supereon
88% of earth history
precambrian division:
Hadean eon
Archean eon
proterozoic eon
followed by Cambrian period of Paleozoic era of Phanerozoic eon

Ancient vs Recent Rocks (there are few older rock deposits available)
major precambrian deposits
gray areas are Phanerozoic
oldest rocks:
issue, Greenland, 3.8 billion years (Ga/By)
approximate locations of Archean cartoons and surrounding Proterozoic regions worldwide, without sedimentary cover

Hadean eon
~4.6 -~4.0 billion years ago
molten earth
lots of extraterrestrial impacts
primitive crust formed
thin early atmosphere of water vapor and volcanic gases
eventual cooling led to massive amounts of condensation and the first global liquid oceans formed
Archean eon
~4.0- ~2.5 billion years ago
surface cooling
tectonic activity began
continents began to form
earliest single-celled life evolved
cyanobacteria (blue-green algae) started producing oxygen by photosynthesis
stromatolites:
layered sedimentary structures
in shallow water
still around today
shark bay in Western Australia
Proterozoic eon
~2.5- ~0.54 bya
great oxidation event
banded iron formations (BIFs)
layered sedimentary rocks with alternating bands of iron oxides
like hematite and magnetite and iron-poor silica (chert)
cyclic/seasonal changes reflected in layers
surface continued to cool
stable supercontinents formed:
Columbia (right) and Rodinia (left)
