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what is a proximate cause
how biological phenomena occur
What is a pre adaptation that’s filling a new function
Exadaptation
what is an ultimate cause
why biological phenomena occur
what is the definition of evolution
inherited changes in properties of populations over the course of generations
what is a theory
hypothesis with enough evidence
what is evidence
results from testing theories
what is a fact
simple confirmable observations
what are the 2 major parts of the origin of species
common descent and descent with modification
what are some aspects of darwins theory
adaptations are passed down through heritable traits within populations, species developed from a common ancestor
what are some aspects of lamarcks theory
adaptations are gained over the lifetime of an individual, species just appeard fully formed
what are the facets of microevolution
natural selection, genetic drift, mutation and gene flow
what is macroevolution
enough microevolution acts that create speciation
what aspect of evolution is shown by a phylogenetic tree
speciation and common descent
what is LUCA
last universal common ancestor
what does monophyletic refer to
when all members of a clade share a common ancestor
what is anagenesis
evolution of traits within a lineage
what is cladogenesis
change of traits that makes 1 lineage into multiple
what are sister groups
two clades that diverged from eachtother
what causes the branch of a phylogenetic tree to rejoin
horizontal gene transfer and hybridization
what does homologous mean
same characteristics inherited from a common ancestor
how can you assume how old characteristics are in a lineage
see how long the branches are on some trees, they can represent how many base pairs have been substituted
what is homoplasy
development of similar traits in multiple lineages (convergent evolution)
what is mosaic evolution
evolution that happens at different rates, due to differing pressure for traits. So different traits can evolve at different rates within a lineage
what is an adaptation
a trait that improves survival and fitness
what is natural selection
selection for a trait that increases fitness within a population
what two things are required for natural selection to occur
a trait must be adaptive and the variation within the trait must be heritable
what traits are used in the “selection for”
traits that increase fitness
what traits are used in the “selection of”
traits that can be located on a nearby locus to the desired trait on the chromosome that can be accidentally selected for when undergoing natural selection
How can we decide if a trait is adpative
complexity: complex traits dont evolve by accident
structure/function: engineering
experiments: can be used to identify traits linked to survival
2 contraints on rapid adaptation and selection
not enough genetic variability and phylogenetic restraints limits on structures
why is adaptation selfish
natural selection can only impact phenotypes it can see are positively aiding the organisms fitness. so cannot adapt for other reasons.
what role does DNA play in inheritance and codons
genome encodes for proteins using codons. Genes are made of three sets of bases which make up a codon.
what is coding DNA
exons, DNA strands used to code genome
what is non coding DNA
introns, cut out and allows for alternate splicing
phenotypes
expressed portion of trait, physical portion
genotype
gene sequence that encode for different phenotypes
segregation
during meiosis only one allele from the parent is included in the gamete
recombination
pair of homologous chromosomes crossing over and swapping genetic material, closer genes to the loci the higher the recombination rate
Harvey Weinburg equillibrium
expected gene frequency from one generation to the next in ideallic population.
what are assumptions made in HWE
infinaite population size, no mutations, no natural selection, no migration and completely random mating. Used as a null model to compare values to.
linkage disequilibrium (D)
non random association between alleles at two loci, will lower genetic variability
linkage equillibrium
if recombination is equally likley between two loci
point mutations
single nucleotide change, in first two postions usually non synonomous and in third usually synonomous
structural mutation
changes in more than one base
whole genome duplication
two diploid gametes fuse together and create offspring with 4 copies of chromosomes
mutation types
deleterious and beneficial
what are mutations important to organism evolution
increases genetic diversity and can help with pre adaptations
pleiotropy
mutation that affects multiple traits, almost always when a mutation impacts phenotype expression
germ line mutations
mutations inherited through gametes
somatic mutations
not inherited but still decrease fitness
what are three types of non genetic inheritance
epigenetics, maternal effects and cultural inheritance
what is absolute fitness (W)
number of offspring an individual will have over their lifetime
relative fitness (w)
compare fitness value among individual by dividing W by a reference
what is selection coefficient (s)
proportional to strength of selection and genetic diversity in a population.
what is genetic correlation
when genes are linked and inherited together when phenotype is selected for
overdominance
Heterozygosity is selected for which aids in maintaining genetic variation
negative frequency dependance
relative fitness is dependant on frequency
underdominance
Heterozygosity decreases fitness which acts to decrease genetic variation
positive frequency dependant selection
selection favoured is whichever is more common in population which also decreases genetic variability
mutation selection balance
natural selection acts to remove deleterious mutations from populations = purifying selection
genetic drift
loss of genetic variation within a population, most common in small popn
how would selection or drift over power eachother
drift is stronger than selection in small population sizes, selection is stronger in larger population sizes and often drives popn to fixation
inbreeding load
fixation of deleterious mutations in small populations which decreases fitness and can lead to extinction
what is the neutral theory of molecular evolution
genetic variation within and among species is primarily due to random genetic drift of mutant alleles that are selectively neutral
discrete traits
traits with clearly outlined character states
quantitative traits
traits that have polygenic inheritance, multiple loci that code for a trait and varies continuously
directional selection of a quantitative trait
moving the mean one way or another
stabilizing selection of a quantitative trait
decreasing the space around the mean, squishing in on ideal values
disruptive selection of a quantitative trait
favours either extreme so increases the overall population variability
how do we measure trait selection strength
relative fitness vs trait value
evolvability (e)
ability for a trait to evolve
what is QTL
quantitative trait loci, regions of genome that affect quantitative traits
what are SNP
single nucleotide polymorphisms, looking for changes in certain DNA bases to understand how phenotypes arose
what does convergent evolution at QTL look like
when a gene encodes for the same thing in multiple lineages
what is local adaptation
traits that thrive in a local population that increase fitness in the area
what is local selection
selection for locally adaptive traits
what is FST
variation between subpopulations, 0: popn have same allele frequency 1: popn are fixed for different alleles
what are the two measures of gene flow
discrete patches and spatially continuous
gene flow and HWE
if huge population shows up with new allele will end up with a lot of homozygotes as the new inhabitants breed with eachother
what is gene swamping
new genes from migrants making homozygotes of a new allele most common which disrupts local variation and selection
How does local selection act on a popn
local selection moves to make populations more different
how does gene flow affect genetic variation within a population
it will act to decrease local genetic variation
what ways can species react to climate change
range shifts, adaptation using pre adaptation possible due to latent variation
BSC
biological species concept, a group of organisms that can interbreed with eachother and are somehow isolated from other groups
what is the phylogenetic species concept
species are distinguishable phylogenetic lineages
what are some pre zygotic reproductive isolation barriers
behavior, environment and sexual, non matching gametes, mechanical
what are some examples of post zygotic reproductive isolation barriers
gene pairing, chromosomal abnormalities
what are DMI
reduction in hybrid fitness, negative interactions between alleles from either parent