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what are the different aspects encompassed by evolutionary biology
-identification and classification of species
-inferring relationships between species
-species adaptations
-genetic variation between species
-how species originated or went extinct
-INVESTIGATION OF THE UNDERLYING MECHANISMS THAT LED TO THE SPECIES OF TODAY
what is the best definition of evolution and who said it
Descent with modification -Darwin (changes in characteristics over generations)
what are the 2 things you need to know to understand how a species evolved
-It’s ancestor
-Modifications that have occurred
what are the mechanisms responsible for evolutionary change
-non-random mating
-mutation
-migration
-genetic drift
-natural selection
what is the definition of natural selection
beneficial alleles increase in frequency due to increased survival and reproductive success of individuals carrying those alleles
are any individuals identical
individuals posess genetic variation
how does the phenotype of an organism change
due to genetic mutations
what effect do genetic mutations have on an individuals fitness
they can increase, decreas or have no effect on fitness
what genetic mutations will increase in a population over time
those that improve an individuals fitness
how are new species, genera or families created
accumulation of advantageous genetic changes over long periods of time
what are the mechanisms that can drive evolution
natural selection, non-random mating, migration, genetic drift
why did the initial HIV drug not work
HIV evolved resistance since it had such a short generation time that a mutation resistant to the drug developed in a short amount of time
how did understanding evolution lead to the modern HIV treatment
the modern multi-drug treatment was developed after it was understood that HIV was evolving resistance to the first drug so the new strategy is just to throw a bunch of drugs at it
are HIV patients worse off taking no medications or only taking some of the treatment
they are worse off taking just some of the treatment since this allows the virus to survive and evolve unchecked
what is methodological naturalism
the strategy of explaining the way the world works using laws of nature
definition of evolution
descent with modification
what is uniformitarianism and who came up with it
charles lyell introduced uniformitarianism which is the school of thought that geological processes operate over long periods of time in a slow gradual manner
what is catastophism
the false school of thought that geological features arose from catastrophes
what was the belief on the origin of life before evolution
spontaneous generation: complex lifeforms arose from non-living matter
how was the theory of spontanepous generation disproven and by who
Francesco Redi with his experiment of raw meat and flies
Who was the first to describe evolutionary change
Erasmus Darwin
what did erasmus darwin describe
all life arose from a single living filament that was modified endlessly into the lifeforms of today and recognised the struggle for existence
who described the principle of progressive development
robert chambers
what was robert chambers recognised for
-new species arise from old species
-diversity of species changes over time
-populations evolve not individuals
-did not explain why diversity occurs
who described the inheritance of acquired characteristics
Jean-Baptiste Lamarck
what did Jean-baptiste lamarck say
new more complex species descend from old less comples species
but he also said that individuals pass down traits they acquired throughout their lives and this was untrue
who described circumstance adaptive law
patrick matthew
what did patrick matthew do
-individuals best suited to their environments will be selected over others
-had most of the idea of evolution but published it in a book about naval timber
who were the 4 predecessors to darwin and what were their theories called
(Charles Lyell (uniformitarianism) Francesco Redi (Disproving spntaneous generation))
Erasmus Darwin-Desribed evolutionary change
Robert Chambers-Progressive development
Jean-Baptiste Lamarck-Inheritance of acquired characteristics
Patrick Matthew-Circumstance adaptive law
What did Charles Darwin do
Published the origin of species which described evolution in a way that hadn’t been before, took a 5 year voyage around the earth’s souther hemisphere
who helped darwin publish his book
Russel Wallace
how did darwin first explain natural selection to his peers
he argued it was like artificial selection and used the dove reference about making a grey dove population int a white one
what are dawins 2 fundamental insights
1:The environment selects on variation in traits of individual organisms
-This answers why species are so well suited to their environments
2:All species descended from one or a few common ancestors
-This answers where species come from
what are the 3 observations that led to the struggle for existence
1:Organisms are very fertile
2:Natural populations remain fairly consistent in size
3:Ressources are limited
what are the 3 observations that led to the discovery of natural selection
1:there is a struggle for existence
2:Individuals within a population Vary
3:Variation is heritable
definition of evolution by natural selection
over many generations accumulation of beneficial traits allows a population to diverge from its original state
is evolution a variational or transformational process
variational
what is a transformational process
ensemble changes because each individual changes (pummeling of sand into dust)
what is a variational process
ensemble changes because something sorts among the variants in the original ensemble (sifting through sand to get dust)
would inheritance of acquired characteristics be a variational or transformational process
transformational
what are the 4pieces of evidence for evolution
1:species change through time (Microevolution)
2:Lineages split and diverge (Speciation)
3:New lifeforms arise from older lifeforms (Macroevolution)
4:All species have arisen from one or a few common ancestors
What is an example of microevolution
The artificial selection of grey pigeons into white pigeons
what is artificial selection
when humans select which individuals reproduce deliberately for a desired outcome
what is the difference between artificial selection and anthropogenic evolution
anthropogenic evolution is an all encompassing term that means evolutionary changes induced by humans whether intentional or not (Ex of unintentional, mountain goats horns getting shorter)
what is the biological species concept
a species consists of groups of actually or potentially interbreeding populations that are reproductively isolated from other groups
what ws the missing trnsitional piece between fish and tetrapods and what is this an example of
tiktaalik, macroevolution
what is homology
shared characteristics as a result of common ancestry
what is biogeography
study of the distribution of species in geographic space and geological time
are species in the same biomes seperated by geographic barriers more or less similar than species in different biomes closer together
less
what are the 3 components of natural selection
1:Variation
2:Inheritance
3:Differential reproductive success
when is a trait heritable
when variation is due to genetic variation
what do evolution researchers study
traits of organisms not the whole organism
do populations evolve or individuals
populations not individuals
when do mutations occur
randomly and independantly of whether they will be favoured by natural selection
how is phenotype determined
by genotype and also the environment
if difference in a trait is due to environmental factors is it heritable
no
what is differential reproductive success
differences in fitness
what is fitness
expected reproductive success of an individual relative to other members of the population
what happened to mice that didn’t match the environmental background compared to ones who did
they were captured more frequently than those that did
does natural selection sort on genotype or phenotype
phenotype
how does natural selection relate to individuals vs a population
it acts on individuals but it’s consequences occur on populations
how do natural selection and evolution interact with allele frequency
natural selection acts on the phenotype but evolution consists of changes in allele frequency
does natural selection lead to a perfect outcome
no, it faces many constraints so never leads to a perfect outcome
what is an adaptation
an inherited trait that increases an organisms fitness and is the result of natural selection for it’s primary function
what is an exaptation
a trait that has one function today but evolved from a trait that served a different function in the past
what is an example of an exaptation
feathers evolved before flight
are most complex traits adaptations or exaptations
most will have undergone many changes with a layering of adaptations and exaptations
what is gene duplication
an entire gene or gene package is duplicated
what is neofunctionalisation
a gene copy created through a copying mistake that acquires a brand new function while the old gene continues to do its old function
how do new hormones/receptors evolve
gene duplication/neofunctionalisation could allow a receptor to mutate and therefore bind to a new hormone
what is gene sharing
a protein recruited to serve a second function somewhere else
what are the 2 ways complex traits evolve
co-opted from previous organs
gradual intermediate steps, each step itself adaptive and serve the same function throughout
what are the constraints on natural selection
short term: lack of sufficient variation
Long term: physical constraints, co-evolution, lack of foresight
why is co-evolution a constraint on natural selection
-can’t optimize phenotype if the thing it’s being optimized for keeps changing
-evolutionary change in one species can effect selective conditions for a second species
-Evolutionary arms race: species invloves each evolve counter measures to the adaptations of the other species
why is lack of foresight a constraint on natural selection
selection can only favour traits that are immediately beneficial, not traits that may be useful in the future
what are the things you need to know to be able to prove a trait is adaptive
determine it’s function
demonstrate that individuls with the trait have a higher fitness than those without
why do mammals tolerate red billed oxpeckers
we think maybe because they serve as an early warning sign, not because they remove ticks, in fact they actually open wounds sometimes
what are the 3 methods of determining a trait’s function
controlled experiment
observational research
comparative methods
what is a controlled experiment
manipulate one component to assess it’s effects on the other components of a system
what is observational research
if a controlled experiment is impractical, careful observation could provide some answers instead
how do comparative methods of research work
look for trends and patterns in evolutionary history by comparing traits of many species, looking for similar traits and functions across multiple species to suggest that it is adaptive
are closely related species statistically independant or not
closely related species may not be statistically indipendant
why is classifying organisms important
helps understand how they’re related to each other
who developed the 8 taxonomic ranks and the binomial nomenclature systems
Carl Linnaeus
what are the 8 taxonomic ranks and in wat order are they organised
Domain→Kingdom→Phylum→Class→Order→Family→Genus→
Species
what are the 2 elements of binomial nomenclature
Genus and Species Epiphet
who is the creator of modern phylogenetic systematics
Willi Hennig
what is phylogenetic systematics and who created it
Invented by Willi Hennig
-Modern approach to classification using evolutionary relationships between organisms
what is the definition of phylogenetics
study of relationships among taxa based on evolutionary history
what is the definition of systematics
study of classification of organisms
what is the goal of phylogenetics
to reconstruct the evolutionary history of a study species and understand their pattern of descent
what are phylogenetic trees built using
traits of interest
what is a character
any observable characteristic or feature of an organism
what is a trait
the specific value of a character
what is the difference between a character and a trait
character is any observable characteristic or feature of an organism while a trait is the specific value of a character
what is the purpose of phylogenetic tree building
to generate a hypothesis
what is a taxon
group of related species placed at the tip of a branch
where are internal nodes on a phylogenetic tree and what do they represent
they are found at the points where branches meet and represent hypothetical common ancestor populations
what do outgroups help you with when creating a phylogenetic tree
they help you root the tree
what is an ancestral trait
a character present in a common ancestor