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What is the closest living relative of animals?
choanoflagellates.
What are the 6 things traits that define/characterize animals?
Cell structure: multicellularity
Mode of nutrition: heterotrophy
Movement: muscle tissue
Reproduction: sexually
Development: all animals possess
How have animal body plans changed since the Cambrian explosion?
They haven’t
What is metazoan organization based on?
Complexity:
Cellular
Tissue
Organ
System
What organism contains no germ layers?
Sponges
What do triploblastic tissues contain?
3 germ layers:
Ectoderm (outer layer)
Mesoderm (middle layer)
Endoderm (inner layer)
What do diploblastic tissues contain?
Ectoderm (outer layer)
Endoderm (inner layer)
What does the ectoderm form?
Epidermis
Hair
Nails
Skin
What does the mesoderm form?
Notochord
Skeletal, cardiac, and smooth muscle
Lining of the coelom
What does the endoderm form?
Lining of the digestive tract
Pharyngeal pouches; tonsils, thymus, and parathyroid
What is radial symmetry and what are its benefits?
Symmetry around a central point
Can interact with the environment from all sides
What symmetry do Cnidarians, Ctenophores (as adults) and some sponges have?
Radial symmetry
What is Bilateral symmetry and what are its benefits?
Symmetry across the sagittal plane; only one way to cut it into equal halves
Well-suited for forward movement and often associated with cephalization
What are the defining features of Protostomes? (bilaterian)
Spiral cleavage
Determinate (mosaic) cleavage
Blastopore becomes mouth
Two major subgroups: Ecdysozoa & Lophotrochozoa)
What are the defining features of Deuterostomes? (bilaterian)
Radial cleavage (cleavage planes are either parallel)
Indeterminate (regulative) cleavage
Blastopore becomes anus
Echinoderms & Chordates
What is an acoelomate body cavity?
no body cavity; space filled with mesodermal cells
only found in triploblastic animals
What is a pseudocoelomate body cavity?
body cavity but not a true coelom as its only partially surrounded by mesoderm
What is an eucoelomate body cavity
Possess a true coelom that’s entirely surrounded by mesoderm
Why do organisms have segmentation?
allows for specialization of body regions
allows functional redundancy (When multiple components perform the same function, so losing one doesn’t cause the function to fail)
What is gamete formation? (1st step of development)
the process by which gametes are formed via gametogenesis
What are the 2 processes of gamete formation?
Spermatogenesis: one spermatocyte (diploid) produces four sperm (haploid)
Oogenesis: One primary oocyte (diploid) produces one egg (haploid) and polar bodies; also produces yolk that provides nutrients to the embryo
What are the 2 things accomplished during fertilization?
Brings haploid genomes together
Activates the egg to begin development; can be internal or external
What are the four steps of fertilization?
Contact; Sperm reaches the jelly coat
Recognition; Species-specific proteins bind to receptors on the egg
Fusion; Sperm and egg membranes fuse; the sperm enters the nucleus
Syngamy; Sperm and egg pronuclei unite forming a diploid zygote
What happens during cleavage?
A large cell divides without growing
How are tissues layers formed via gastrulation?
Invagination; the inward folding of the blastula
What are the four ways to distinguish protostomes and deuterostomes
Cleavage (spiral vs. radial)
Cell fate (determinate vs. indeterminate)
Blastopore fate (mouth first vs. anus first)
Coelom formation (Schizocoely (splitting) vs. Enterocoely (pouching/pinching)
What level of organization do sponges have?
Cellular (no tissues, no organs)
What is a Pinacocyte?
flat sponge cell that forms the outer covering of the sponge
What is a Porocyte?
tube shaped sponge cell that controls water flow through ostia
What is an Archeocyte?
an amoeba-like cell that delivers nutrients to cells, and differentiates into other cell types
What is a Sclerocyte?
a cell that secretes spicules
What is a Choanocyte?
a flagellated cell that generates water current and filters food particles from water
What is the path of water through a sponge?
Comes in through the ostia (small pores)
Flows through incurrent canals
Water enters choanocyte chambers where it is filtered
Carried toward the spongocoel (central cavity) via excurrent canals
Water leaves through oscula (large, visible openings)
How do choanocytes generate current and capture food?
a singular flagellum lined with microvilli (tiny, finger-like projections of the cell membrane; increasing surface area)
What are the features of the asconoid canal system?
Single, large osculum
Water comes in through the ostia into the spongocoel, which is lined with choanocytes
What are the features of the syconoid canal system?
Single, large osculum
Spongocoel lining is folded back to make radial canals which are lined with choanocytes
Choanocytes in radial canals rather than spongocoel
What are the features of the leuconoid system?
No spongocoel
Complex folding with choanocyte-lined chambers
multiple oscula
Most sponges have this system
What are the defining features of Calcispongiae?
Calcium carbonate spicules
Mostly small and shallow water; all 3 canal systems
What are the defining features of Hexactinellida (glass sponges)?
6-rayed silica spicules
Mostly deep sea
What are the defining features of Demospongiae?
Silica spicules (not 6-rayed), spongin, or both
Most living species; freshwater & bath sponges; leuconoid
What are the defining features of Homoscleromorpha?
Small silica spicules, or none
have a basement membrane; the closest thing to epithelium
How do sponges reproduce?
Both sexual and asexual
Asexual via bud formation/fragments
Sexual: hermaphroditic; fertilization
What are the four things that have allowed sponges to persist for so long?
Spicules; skeletal elements that make sponges hard and unpleasant to eat
Chemical defenses; toxic/foul-tasting compounds that deter predators & competitors
Regeneration
Microbial partners; symbiotic bacteria take u; a share os sponge biomass and help produce defensive chemicals
What is the defining trait of Lophotrochozoans?
have either a permanent lophophore feeding apparatus or a trochophore larvae
What are the defining traits of Phylum Platyhelminthes?
1st appearance of bilateral symmetry; Flatworms
Triploblastic; acoelomate body cavity and a blind (one-way) gut
What is the importance of Platyhelminthes having a flattened body?
Higher surface area to volume ratio
Since they have no respiratory or circulatory system, it allows cells to get nutrients & oxygen via simple diffusion
Is there a nervous system present in Platyhelminthes? Are they able to excrete waste and osmoregulate?
They have a nervous system, one of the earliest appearances of cephalization, and eye spots (Ocelli)
Excretory/osmoregulatory system works via flame cells (protonephridia)
What are the defining features of Class Turbellaria?
Mostly free-living
Muscles and cilia for movement
Some gliding over mucous from adhesive glands
What are the defining features of Class Monogea?
Ectoparasites, mainly of fish
Single host (direct life cycle)
Attach to gills and feed on body fluids
Only damaging in crowded conditions
What are the defining features of Class Trematoda?
All parasitic flukes (endoparasitic in vertebrates)
Suckers and hooks for adhesion; minimal sense organs
Indirect life cycle with a intermediate and definitive hos
Important parasites of humans and domestic animals
What are the defining features of Class Cestoda (tapeworms)?
All endoparasites, usually in the vertebrate gut
No mouth or digestive tract
Hooks and suckers for attachment (Scolex)
Usually 2 hosts
What are the defining features of Phylum Bryozoa?
Moss animals; mostly aquatic sessile organisms; Both freshwater and marine (shallow)
Bilateral symmetry; radial & mosaic (determinate) cleavage
Coelomate
How do Bryozoans feed and reproduce?
Filter feeders
Asexual: budding
Freshwater; Statoblasts: hard-shelled asexual capsules that survive freezing and drying
Sexual; hermaphroditic
What are the features shared by radial animals?
radial symmetry
diploblastic; contain tissues
most have blind guts; ctenophores exception
What are the morphological differences between the polyp and medusa body forms?
The polyp is the sessile, tube-like form
The medusa is the free swimming, umbrella-shaped form
What are advantages of having a life cycle with both polyp and medusa life forms?
Division of labor; the polyp can stay put in a favorable environment, feed and perform asexual reproduction while the medusae performs sexual reproduction, allowing for better dispersal and the opportunity to get to a better habitat?
Where do you find the cnidocytes and how do cnidocytes and nematocysts work in prey capture and defense?
Cnidocytes are specialized stinging cells that are found in the epidermis
When the cnidocil (modified cilium) detects touch or pressure, the operculum (lid) opens, leading to the cnidocytes discharging the nematocyst
What are the defining features of Class Hydrozoa?
Commonly colonial; organized as a branching stalk of polyps
Medusa stage present; typically includes polyps and medusae
Asexual via budding; Sexual fertilization with medusa
What is the life cycle of Hydrozoans?
A polyp colony asexually buds medusae. Medusae release gametes; fertilization produces a planula larva, which settles and becomes a new polyp colony.
What are Gastrozooids, Gonozooids, and Dactylozooids?
Gastrozooid: feeding polyp
Gonozooid: reproduction polyp
Dactylozooid: defense and prey capture polyp
What are the defining features of Class Scyphozoa?
True jellyfish, mostly floating
Medusa dominant
Move by pulsing the bell
What is the life cycle of a scyphozoan (true jellyfish)?
Adult medusae release gametes, and fertilization produces a ciliated planula larva. The planula becomes a small polyp (scyphistoma) which buds and releases juvenile medusae called ephyrae. Ephyrae grow into adult medusae.
What are the defining features of Class Cubozoa?
Box jellies with bells
Dominant form: medusa
Fast swimmers; can produce fatal stings
WHat is the life cycle of Cubozoans?
Adult medusae release gametes, producing a planula larva. The planula settles as a small polyp, which can bud more polyps. Each polyp then metamorphoses entirely into a single juvenile medusa, which grows into an adult.
What are the defining features of Class Anthozoa?
All polyps, no medusa stage
Sea anemones. corals
Calcium carbonate skeletons
Have zooxanthellae (symbiotic algae) living in their tissues
What is the symbiotic relationship between anthozoans and zooxanthellae?
Zooxanthellae supply sugars and other nutrients and boost calcification. The coral provides shelter, CO₂, and nitrogen wastes.
How do anthozoans reproduce?
Sexually: gametes are released by broadcast spawning (external fertilization) or brooded internally, forming a planula larva that settles as a new polyp. Asexually: budding, fragmentation, longitudinal fission, and pedal laceration.
What are the defining features of Ctenophores?
Comb-like plates (cteno)
Colloblasts (specialized glue cells found on tentacles)
Complete gut
How do ctenophores reproduce?
Mostly sexually. They are hermaphrodites whose gametes are released through the mouth or body wall for external fertilization. Development is direct(no polyp or medusa stage). Asexual reproduction is rare, though some can regenerate or fragment.
What are the 6 defining traits of molluscs?
Radula (rasping feeding organ)
Mantle (secretes shell)
Muscular foot for locomotion
Complete respiratory system with gills or lungs
Circulatory system with a heart
Ciliary tracts
What are the 3 parts of mollusks body plan?
Head-foot (contains feeding, locomotory, and sensory organs)
Visceral mass (contains digestive, circulatory, respiratory, and reproductive organs)
Mantle (secretes shell)
What are the mollusks embryonic features?
Triploblastic (3 germ layers)
Protostomes (1st mouth)
Spiral & Mosaic Cleavage
Coelomates (via schizocoely)
Most molluscs have an open circulatory system. What does that mean and how does it compare to open circulatory systems?
An open circulatory system means blood isn’t always confined to vessels
In a closed circulatory system, blood stays confined to vessels
What is the process of torsion and what are the pros and cons?
The anus comes toward the mouth creating the mantle cavity
Pros: Protection
Cons: Fouling
What is the process of coiling and what are the pros and cons?
Coiling is the winding of the shell into a spiral
Pros: Mobility, Less Stress due to weight, Protection
Cons: Asymmetry (leading to organisms losing organs on one side to balance), Imbalance, Limited flexibilty
What are the defining traits of Gastropoda?
If present, shell is in single piece (except nudibranchs)
Undergo torsion in larval stage (veliger)
Undergo coiling of shell and visceral mass
How do Gastropods feed and perform respiration?
Feed with a radula
Some use cilia or mucous for food capture
Usually one efficient gill or lung
How do Gastropods reproduce?
Usually dioecious, some hermaphroditic
Internal fertilization
Offspring encapsulated in egg cases/capsules
What are the defining traits of nudibranchs?
Show detorsion; no shell
Warning coloration
Get defenses such as nematocysts or toxins from cnidarians and sponges
What are the defining features of Class Bivalvia?
2 shells (valves)
Most are filter feeders (no radula)
No head (reduced cephalization)
Most marine
Open circulatory, excretory, and respiratory systems present
How do Bivalves open and close?
Dorsal ligament (opening)
Adductor muscles (closing)
How do Bivalves move?
Foot is extended between valves, blood is pumped, causing it to swell and act as an anchor
Some bivalves are sessile
How do Bivalves reproduce?
Most dioecious; some hermaphroditic and some oysters can chnage sex
Broadcast spawners (eggs & sperm released into water); one oyster can release 1 million eggs
External fertilization
What are the defining traits of Class Cephalopoda?
All marine & carnivorous
Shell reduced; absent in Octopus
Highly modified foot, moved toward head
Locomotion via jet propulsion through siphon
What is a siphon?
Tube-like structure near the head
Modified mantle cavity opening; main outlet for water
Central to jet propulsion
How are Cephalopods composed?
No cilia on gills
Large brain; complex eyes with excellent eyesight (colorblindO
Ink gland for defense
How do Cephalopods reproduce?
Dioecious; courtship via colorful displays
Male’s modified arm (hectocotylus) transfers sperm packets (spermatophores)
Large, yolk-rich eggs
Direct development
Semelparous (breed once and die)
How do cephalopods communicate?
Chromatophores (pigment cells in the skin)
Expand and contract to induce color changes
What are the defining traits of Annelida?
1st appearance of segmentation/metamerism
Coelom segmented w/septa (thin will of tissue)
Setae (bristle-like structures composed of chitin) for locomotion or an anchor
Complete digestive system, nervous system, excretion, closed circulation, respiratory system
Trochophore as ancestral larval form
What are the traits of Annelid embryonic development?
Triploblastic
Protostomes
What unites Clade Ctillata?
All possess a reproductive structure called the clitellum
Ring of secretory cells around the body
What is the difference between oligochaetes and leeches?
The clitellum is always visible in oligochaetes; present only during the reproductive seasons in leeches