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What has been a feature of coastal oceans for nearly as long as there has been life on earth?
reefs
What has been a feature of reefs since the beginning of time?
photosynthesis
What has changed many times?
the types of organisms that have built reefs?
What has caused the types of organisms who have built reefs to change?
mass extinctions, evolution of community ecology, changes in sea levels, tectonics, and changes in the ocean geochemistry
what is the study of the chemical composition of sea water, ocean sediments, and the marine crust, as well as the chemical processes and cycles that shape them
ocean geochemistry
What period of reefs occurred 3.4-0.54 billion years ago?
Precambrian reefs
how were precambrian reefs formed before multicellular organisms?
by photosynthetic cyanobacteria
What did the photosynthetic cyanobacteria produce?
stromatolites
How are stromatolites formed?
built by cyanobacteria through microbial mats trapping sand silt and minerals which are cemented and then the microbes migrate upward to stay in the sunlight and the process happens again
What are stomatolites?
stromatolites are layered rock like sedimentary structures
What period of reefs occurred 541-443 million years ago?
Early Paleozoic reefs
What built early paleozoic reefs?
Archaeocythid sponges
Did the archaeocyathic sponges have photosynthetic symbionts?
they may have
What was a trait of early paleozoic reefs?
many growth forms
What led to the extinction of early paleozoic reefs?
sea level rise, high co2 and global warming
What happened during the early paleozoic reefs era?
the Great Ordovician Biodiversification Event
What was the Great Ordovician Biodiversification event?
a diversification of reef-building biota that was biologically driven rather than being driven by environmental change
What organisms became reef builders after the Great Ordovician Biodiversification Event?
bryozoans, sponges (lithistid sponges and stomatoporoids), brachipods, and rugose corals
What appeared during this time but didn’t start building reefs until the end of the Ordovician?
Tabulate corals
What corals are solitary and colonial, have well-defined septa, well-defined tabula, are often horn-shaped and have calcite skeletons?
Rugose corals
What corals are nearly always colonial, have poorly defined septa, well-defined tabula, and calcite skeletons?
Tabulate corals
What are the main differences between rugose and tabulate corals?
rugose can be both solitary and colonial and have well defined septa, while tabulate corals are nearly always colonial and have poorly defined septa.
What appears during the Mid-Paleozoic?
the first coral reef ecosystems
What geologic era was 443-359 million years ago?
the Mid-Paleozoic
What association dominated on coral reef ecosystems during the Mid-Paleozoic?
coral-stromotoporoid-algae association
What geologic era had a super greenhouse Earth?
Mid-Paleozoic
what is a supergreenhouse Earth?
An extreme climate state in earth’s geologic history characterized by drastically elevated greenhouse gas concentrations, severely elevated temperatures, and near-total or complete absence of polar ice sheets
What latitudes did reefs grow at in the Mid-Paleozoic?
55 degrees latitude
Were reefs much larger in the Mid-Paleozoic than today?
yes reefs were much larger
What wiped the reefs out in the Mid-Paleozoic?
the Devonian mass extinction
What is likely to have caused the Devonian mass extinction?
climate change
Which geologic era contained reefs that has large colonies of tabulate and rugose corals, crinoids, bryozoans, and brachipods?
the middle silurian during the mid-late paleozoic
Which geologic era occurred 359-252 million years ago?
the Late Paleozoic
What happened to reefs during the late paleozoic
reefs diversified but struggled until the permian extinction?
Did the late paleozoic have calcite or aragonite seas?
aragonite seas
What was the largest mass extinction to hit marine ecosystems?
the permian extinction
What happened during the permian extinction to corals?
rugose and tabulate corals were wiped out alongside stromatoporoids and the reefs they built
What was likely to have caused the permian extinction?
volcanism contributing CO2 to the atmosphere and oceans, dramatic warming, acidification, and anoxia
what is anoxia?
Total absence of dissolved oxygen in water, soil, or sediment
What geologic era occurred 252-66 million years ago?
the Mesozoic
What did we see for reefs during the mesozoic?
the first modern reefs
How long were the reefs gone after the permian extinction?
about 8 million years
What was atmospheric CO2 like during the Mesozoic?
huge fluctuation in atmospheric CO2
What did this huge fluctuation of atmospheric CO2 in the Mesozoic cause?
lots of little diversifications and extinctions?
When do the first scleractinians arrive during the Mezoic?
during the middle triassic
When do the scleractinians that arrive during the Mesozoic start building reef?
during the Late Triassic
During the jurassic-cretaceous transition what appear and build reefs with corals?
rudists
How long do corals and rudist bivalves build reefs for?
millions of years
What happened with rudists in the early Cretaceous during the Mesozoic?
they replace corals mysteriously
What type of seas did the cretaceous period in the Mesozoic have?
calcite
What were rudist bivalves?
mollusks
did rudist bivalves have algal symbionts?
yes
what types of shells did rudists have?
both calcite and aragonite
What happened to both rudists and reefs at the end of the Cretaceous during the Mesozoic?
they were wiped out by a meteor impact
What period was 66-0 million years ago?
the Cenozoic
How many genera of scleractinians survived into the Cenozoic after the extinction at the end of the Cretaceous period?
11
What was it like for reefs during the Cenozoic?
they struggled for a while on a greenhouse earth
What period do we eventually see reefs as we know them
cenozoic
What is the difference between aragonite and calcite?
calcium carbonate crystals come in several forms and different levels of magnesium leads to either calcite or aragonite
What does low magnesium lead to?
calcite
What does high magnesium lead to?
aragonite
Which is stronger but less stable?
aragonite
what type of skeletons do modern corals build?
aragonite
what types of skeletons do mollusks tend to build?
calcite
What causes one crystal to be favored over another?
ocean chemistry
What do icehouse seas favor?
aragonite
what do greenhouse seas favor?
calcite
What happens are the ocean warms?
calcite is favored
why does volcanism favor calcite
because it eats up magnesium and with less magnesium you have calcite
What changes with the environmental fluctuations of greenhouse-icehouse and calcite versus aragonite?
the winning reef builders shift