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nomenclature of hybrids
new genus is a combined name of two genera, X is used to indicate the cross
three domains of life
eukarya, bacteria, archaea
endosymbiont theory
theory that proposes that mitochondria and chloroplasts were once individual prokaryotic organisms that eventually integrated to become one eukaryotic cell
evidence of endosymbiont theory
inner membranes similar to plasma membranes of prokaryotes, organelles create their own DNA, division and DNA structure are similar in organelles and prokaryotes, their ribosomes are more similar to prokaryotic
angiosperms appeared about..
360 million years ago
origin of plants
470 mya
chlorophytes
division of green algae
contain chloroplasts, cell walls, and use starch as storage for photosynthates
ancestor of land plants
green algae
first plants to grow on land
non-vascular plants (ex. moss, liverworts) around 475 mya
move to land involved
creation of soils, development of a cuticle
bryophytes
reproduce by spores, requiring water
have rudimentary leaves, roots, or stems
grow in moist, shady areas
sporophyte generation (diploid 2n)
produce haploid spores via meiosis
spores grow into gametophytes
gametophyte generation (haploid n)
produce haploid gametes by mitosis
fusion of gametes (fertilization) produces diploid sporophyte
mitosis
begins with one haploid or diploid cell, produces two identical daughter cells (two haploid or diploid)
meiosis
first meiotic division produces two haploid, second produces four haploid
seedless vascular plants
first vascular plants, appeared around 425 mya
vascular system
xylem and phloem carry nutrients around plant
allowed plants to grow tall
xylem
carries water up from roots and transports
phloem
carries sugars made via photosynthesis in leaves and transports
angiosperms
flowering plants producing seeds within protective fruit/ovary
monocots
single cotyledon, parallel veined leaves
eudicots
4 or 5 merous flowers, branching-veined
“lower plants”
mosses, ferns
“higher plants”
angiosperms, gymnosperms
reproductive shoot of angiosperms
flower
carpel
consists of ovary at the base and style leading up to a stigma where pollen is recieved
flower consists of…
speals, petals, stamens, carpels
stamen
consists of stalk, with anther where pollen is produced
methods of pollen attraction in flowers
colour, nectar, odor, mimicry/deception
pollinator reward
pollen and nectar
pollen
produced by male parts of seed plants for reproduction, eaten by animals (contains nutrients)
nectar
sugar-rich liquid produced by plants, is relied on by pollinators (ex. butterflies) for all their nutrition for several months
plant trickery - mimicry
flowers can mimic insect shapes, inducing a response from insects (ex. reproduction, fight), and results in the uptake of pollen and transferration
examples of pollinators
bees, butterflies, bats, birds
characteristics of plants pollinated by bats
open at night, dull colours, positioned below foliage, exude musty odor, large and tough with lots of pollen/nectar
characteristics of plants pollinated by birds
red/orange in colour, deep tubes without landing platform, often horizontal, have lots of nectar but emit no odor
co-evolution
plants and the animals that pollinate them evolved at the same time and in ways that benefit the other
wind-pollinated flowers
small, colourless, odorless, no nectar, no petals (or very small petals), dangle and wave in the open, produce more pollen grains per ovule compared to animal-pollinated
wind-carried pollen grains
small, smooth, and dry, 20-60 micrometers in diameter, shape is frisbee-like
angiosperm life cycle
sporophyte-dominant, heterosporous - producing microspores and megaspores
microspores
generate pollen grains as the male gametophytes
megaspores
form an ovule that contains female gametophytes
anthers
pollen-producing part at top of a flower’s stamen (male component)
made up of four lobes called pollen sacs
pollen sacs
found within anthers, containing a mass of dividing cells called microsporocytes
microsporocytes
within pollen sacs in anthers
cells that divide via meiosis forming four haploid microspores
microspores
generated by microsporocytes in pollen sacs
each microspore divides via mitosis forming two-celled pollen grain. role is to produce sperm cells for ferilization
pollen tube
produced when germination of a pollen grain occurs, grows through stigma and style and enters ovary
double fertilization
two sperm cells created by the generative cell pass through a synergid cell - one fuses with egg the the other with central cell. fusion of sperm and egg cell forms a diploid zygote which grows into an embryo, while fusion of sperm nucleus with polar nuclei forms triploid nucleus, which divides to become a reserve tissue called the endosperm
types of angiosperms
complete/perfect, incomplete/imperfect, inflorscence
complete/perfect angiosperms
contain all 4 floral organs/whorls (ex. apple)
incomplete/imperfect angiosperms
lack one or more floral organs/whorls such as stamens or carpels
inflorescence angiosperms
clusters of flowers (ex. wild carrot)
monoecious plants
male and female flowers are separate, but carried on the same plant. developmental timings are asynchronous
self-pollination or selfing
transfer of pollen from anther to stigma in the same flower, and transfer of pollen from one flower to another on the same plant
cross-pollination or outcrossing
involves transfer of pollen from one genetically distinct plant to the stigma of another (increases genetic diversity)
dioecious plants
plants have either male or female flowers
s-genes
recognition of self is based on this - genes for self-incompatibility
there can be many alleles of this
gemtophytic incompatibility
if a pollen grain has an allele that matches an allele of the stigma it lands on, the pollen tube fails to germinate or fails to grow through the style to the ovary
sporophytic incompatibility
the pollen phenotype is determined by the diploid genotype of the plant on which it was produced because proteins corresponding to both alleles of the pollen parent are deposited into the pollen coat
self-fertilization advantages
widespread propagation of the same adapted genotype
no need for wind or pollinators
little metabolic cost (little wasted pollen, no cost to produce nectar)
self-fertilization disadvantages
restricted opportunity to create new genetic combinations
inbreeding depression
loss of vigour
fruit
enlarged ovary that contains seeds
seed
embryo and nutrient source surrounded by a protective coat
flower
reproductive structure that produced gametes, protects and nourishes megaspore gamete, captures male gametes, nourishes embryos, develops seeds/fruits
simple fruit
fleshy or dry, dry can either be dehiscent or indehiscent
develops from a single carpel or several fused carpels of one flower
aggregate fruit
develops from many separate carpels of one flower
accessory fruit
develops from tissues other than ovary (ex. strawberry)
pathenocarpic
fruits that develop without fertilization, seedless (ex. grapes, banana, pineapple)
stimulative parthenocarpy
pollination or other stimulation is required for parthenocarpy
vegetative parthenocarpy
exhibited by plants that dont require pollination or other stimulation to produce parthenocarpic fruit
fruit and seed dispersal
plants life depends on finding fertile ground
to prosper seeds must be widely dispersed
use of biotic or abiotic dispersal agents
evolution modifications for wind dispersal
winged/plumed fruits
ballistic dispersal by violent dehiscence of pericarp
evolution modifications for water dispersal
buoyancy in seeds, surviving for months/years at sea
water pushes seeds against rocks, scraping the coat to remove dormancy
mechanical dormancy
scarification allows seeds to become active (stopping dormancy)
fruit as dispersal aids
stored food in fruit is not utilized by dormant seeds
animal vectors are used - many seeds in fruits can survive passage through animal digestive tracts
dispersal by animals
seeds get caught in fur, passed through digestive tracts, hoarding animals move seeds/fruits underground
seed dormancy
increases chances that germination will occur at a time and place most advantageous to the seedling
environmental cues are often required to break dormancy
physical dormancy
impermeable seed coat, mechanical dormancy
broken by passage through digestive tracts, mechanical nicking, stratification
components of dicot
radicle in centre, surrounded by two cotyledons
radicle has an epicotyl on the top and hypocotyl below
seed coat surrounding seed
components of monocot
one cotyledon (called scutellum)
rudimentary shoot is sheathed in coleoptile, coleorhiza covers young root
pericarp is fused with the seed coat - inside is the endosperm, radicle with epicotyl and hypocotyl, scutellum, coleoptile, and coleorhiza
seed germination
first step of growth of the embryo
after imbibition, enzymes activated and rapidly released to digest stored food into smaller molecules - which are transported and converted into energy needed for growth
germination - what it looks like
appearence of a radicle and shoot from the seed
emergence - what it looks like
appearance of a seedling shoot above the soil surface
epigeal
cotyledons emerge above the ground
hypogeal
cotyledons remain in the soil