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What do echinoderm fossils and larvae indicate?
That they are bilaterally symmetrical and pentaradial symmetry is dervied
What type of symmerty can be limiting for free-moving animals?
Radial symmetry
Where are echinoderms abundant?
In marine habitats
Why are echinoderms unique?
They possess features not seen in other phyla
How did hemichordates develop to differ from echinoderms?
They developed buccal diverticulum and mesocoelomic ducts
What superphylum do echinoderms and hemichordates belong to?
Deuterostomes
What trait were present in stem echinoderms but lost in modern echinoderms?
Gill slits
Are echinoderms vertebrates or invertebrates?
Invertebrates
Are hemichordates vertebrates or invertebrates?
Invertebrates
What does the clade ambulacraria contain?
Echinoderms and hemichordates
What are examples of echinoderms?
Sea stars and brittle stars
What are examples of hemichordates?
Acorn worms and pterobranchs
What do members of the clade ambulacraria share?
Deuterstome features, three-part coelom, similar larval forms, and axial (specialized filtering) complex
What combination of characteristics are found in echinoderms but no other phylum?
A calcareous endoskeleton, large plates/small ossicles, a water-vascular system, pedicellariae, dermal brachiae, and pentaradial symmetry in adults
What type of symmetry do adult echinoderms have?
Pentaradial symmetry
Were most early forms of echinoderms free-moving or sessile?
Sessile with radial symmetry
What class of echinoderms still has sessile forms?
Crinodiea
Are most modern echinoderms sessile or free moving?
Free-moving
What are ancestral deuterostomes envisioned as?
Benthic filter-feeder ingesting particles via ciliary water movement
What are the two forms of echinodermata?
Yanjiahella and ctenoimbricata
What are characteristics of the Yanjiahella form of echinoderms?
It is a bilateral, two-armed form
What are characteristics of the Ctenoimbricata form of echinoderms?
It is a benthic, bilateral forn with a ventral mouth
While all echinoderms are marine organisms, they have a limited ability to do what?
Osmoregulate
Where are most adult echinoderms?
At low points of the sea (benthic)
While some echinoderms are particle feeders, many are what?
Predators
What phylum do regular and irregular sea urchins belong to?
Echinoderms
What do regular sea urchins feed on?
Algae
What do irregular sea urchins feed on?
They are deposit feeders
What phylum do crinoids belong to?
Echinoderms
What do crinoids feed on?
Plankton and suspended particles
What echinoderms are the most active?
Brittle stars
How do brittle stars feed?
Thy are scavengers, browsers, and deposit or filter feeders
What do some brittle stars have a commensalism relationship with?
Sponges
What type of axis do sea cucumbers have?
An extended oral-aboral axis
What is the sea cucumbers primary feeding method?
Suspension or deposit feeding
What class of echinoderms are sea stars?
Asteroidea
Where are sea stars commonly found?
Along shorelines, rocks, muddy/sandy bottoms, or coral reefs
Approximately how many living species of sea stars are there?
1500
What size are sea stars?
Range from a cm to a meter across
What color are sea stars?
Often bright colored
What are the external features of asteroids (sea stars)?
A central disc with 5+ tapering and extending arms
What is the oral surface of an asteriod (sea stars)?
The surface with the mouth
What is on the aboral surface of an asteriod (sea stars)?
The surface opposite of the mouth
What is the ambulacrum of an asteroid (sea star)?
Region from the mouth to the tip of each arm
What borders the ambulacral groove of an asteroid (sea star)?
Rows of tube feet
What borders the rows of tube feet of an asteroid (sea star)?
Moveable spines
What are at the end of arm of an asteroid (sea star)?
Sensory tentacles
What is the circular madreporite of an asteroid (sea star)?
A porous plate that allows water to enter the water-vascular system
Where is the circular madreporite of an asteroid (sea star) located?
On the aboral surface
Where is the radial nerve of an asteroid (sea star) located?
In the center of the ambulacral grooves
What is under the radial nerve of an asteroid (sea star)?
The extension of the coelom
What is the extension of the coelom of an asteroid (sea star)?
The radial canal of the water-vascular system
Ossicles/dermal tissues that cover the structure of asteroids (sea stars) are present in all classes expect what?
Crinoids
What is the usual texture of the aboral surface of an asteroid (sea star)?
Rough and spiny
What do many asteroid (sea star) species have that keep the body free of debris?
Pincer-like pedicellariae
What are papulae (dermal branchiae or skin gills) of an asteroid (sea star)?
Soft projections between ossicles for gas exchange
What penetrates between the ossicles of an asteroid (sea star)?
Tube feet (podia)
What type of pedicellaria of an asteroid (sea star) are associated with muscles?
Forceps-type pedicellaria
What is under the epidermis of an asteroid (sea star)?
The mesodermal endoskeleton
What is the mesodermal endoskeleton of an asteroid (sea star) made of?
Small calcareous plate (ossicles)
What binds the mesodermal endoskeletonWossicles of an asteroid (sea star) together?
Connective tissue called catch collagen
What is the stereom of an asteroid (sea star)?
The porous, mesh-like structure of the ossicles with spaces filled with fibers and dermal cells
What gives rise to several adult structures of adult asteroid (sea star) structures?
Larval coelomic compartments
What is the coelom of an asteroid (sea star)?
A spacious fluid-filled body cavity
What circulates fluid through the coelom to the papulae of an asteroid (sea star)?
The ciliated peritoneal lining
How do asteroids (sea stars) remove gases and waste?
Throgh diffusion across the papulae and tube feet
What gas do asteroids (sea stars) primarily diffuse?
Ammonia
What organs do asteroids (sea stars) replace with diffusion?
Kidneys and lungs
What system is a coelomic compartment that is unique to echinoderms?
The water-vascular system
What does the water-vascular system of an asteroid (sea star) consist of?
A system of canals and tube feet with dermal ossicles the form a hydraulic system
What are the functions of water-vascular system of an asteroid (sea star)?
Locomotion, food-gathering, respirattion, and excretion
What does the madreporite of an asteroid (sea star) lead to?
The stone canal → the ring canal that encircles the mouth
What extends from the ring canal of an asteroid (sea star)?
Radial canals that extend into each ray
What functions as fluid reservoirs in the water-vascular system of an asteroid (sea star)?
Polian vesicles that can attach to the ring canal
What connects the radial canal to the tube feet (podia) of an asteroid (sea star)?
Small lateral canals, each with a one-way valve
What does the inner end of a tube foot of an asteroid (sea star) consist of?
An ampulla that lies within the coelom
What does the outer end of a tube foot of an asteroid (sea star) consist of?
A sucker
What do the ampulla muscles of the water-vascular system of asteroids (sea stars) do?
Contract and force extension and fluid into podium
What forces fluid back into the ampulla from the podium of an asteroid (sea star)?
Contraction and retraction of the tube foots longitudinal muscle
What creates suction for locomotion of an asteroid (sea star)?
When the small muscles in the end of the tube foot raise in the middle section
How much pull can the suction of tube foot of an asteroid (sea star) create?
0.25 - 0.30 newtons
What allows an asteroid (sea star) to move vertical surfaces?
Coordinated action of all or many tube feet
How is the locomotion of an asteroid (sea star) different on soft surfaces?
Suckers are ineffective, causing tube feet to operate like legs
What does the mouth of an asteroid (sea star) lead to?
A two-part stomach in the central disc
What are the two parts of the stomach of an asteroid (sea star)?
The larger lower cardiac stomach and the smaller upper pyloric stomach
Which stomach of an asteroid (sea star) can be everted through the mouth during feeding?
The large lower cardiac stomach
What is the upper pyloric stomach of an asteroid (sea star) connected to?
Pyloric ceca (digestive glands) in each arms
Where does digestion occur in an asteroid (sea star)?
Mostly extracellularly in the pyloric ceca
Do all asteroids (sea stars) have an intestine and anus?
No, some species lack both
What do asteroids (sea stars) feed on?
Invertebrates, vertebrates, and particles
What species of asteroids (sea stars) prey on mussels and oysters?
Asterias
What species of asteroids (sea stars) is damaging the coral reef in the Pacific Ocean?
Crown-of-thorns starfish
How are crown-of-thorns starfish damaging the coral reef?
They feed on coral polyps
How do the diets of crown-of-thorns starfish change based on their population size?
Low population, they eat the faster-growing corals. High population, they eat all corals
What causes the population of crown-of-thorn to increase?
Nutrient run off increases plankton population. Plankton are food for larval sea stars
What is the hemal system of an asteroid (sea star)?
A system of tissue stands enclosing perihemal channels
What does the hemal system of an asteroid (sea star) play a role in?
Distributing nutrients
Is the hemal system of an asteroid (sea star) developed fully?
No, it is not developed well
What do echinoderms lack in their nervous system?
A brain and distinct ganglia
How many subsystems does the nervous system of echinoderms have?
Three systems that each have a ring and radial nerves, and occur at different levels in the disc and arms