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Cotransporter
Protein that moves two substances together; one moving down its gradient can power movement of another against its gradient.
Plant
Multicellular eukaryote; mostly autotrophic; chlorophyll; cellulose cell walls; alternation of generations.
Photoautotroph
Organism that uses light for energy and makes organic carbon from inorganic carbon.
Phototroph
Organism that obtains energy from sunlight.
Chemotroph
Organism that obtains energy from chemical compounds.
Autotroph
Organism that produces its own organic compounds from inorganic carbon.
Heterotroph
Organism that obtains organic carbon from other organisms or compounds.
Nutrient
Substance required for biosynthesis and normal growth.
Macronutrients in plants
Elements required in large amounts; includes CHONPS plus potassium, calcium, and magnesium.
Micronutrients in plants
Elements required in trace amounts, including iron, manganese, boron, molybdenum, copper, zinc, chlorine, nickel, cobalt, sodium, and silicon.
CHONPS
Carbon, hydrogen, oxygen, nitrogen, phosphorus, and sulfur.
Carbon function in plants
Backbone of carbohydrates, proteins, fats, and nucleic acids.
Nitrogen function in plants
Needed for proteins, nucleic acids, and some vitamins.
Phosphorus function in plants
Important in nucleotides and ATP.
Sulfur function in plants
Found in some amino acids, coenzymes, and vitamins.
Gaseous nutrient sources
Elements found as gases in the atmosphere or oceans; examples include oxygen, carbon, hydrogen, and nitrogen.
Sedimentary nutrient sources
Mineral salts released by weathering of rocks; examples include phosphorus, calcium, and some sulfur.
Soil
Mixture of minerals, organic materials, living organisms, air, and water covering land.
Parent material
Inorganic mineral matter in soil; often about 40-45% in the lecture's overview.
Humus
Organic matter in soil; about 5% in the lecture's overview.
Soil horizons
Distinct vertical layers created during soil formation.
Soil profile
Vertical slice showing the different soil horizons.
Weathering
Physical or chemical breakdown of rock into smaller particles.
Sand particle size
0.063-2.0 mm in diameter.
Silt particle size
0.004-0.063 mm in diameter.
Clay particle size
Less than 0.004 mm in diameter.
Soil texture
Relative proportion of sand, silt, and clay in a soil sample.
Loam
Soil with a reasonable mix of sand, silt, and clay; retains nutrients and some water while draining adequately.
Leaching
Loss of dissolved nutrients as water moves through soil.
Soil aeration
Availability and movement of air, including oxygen, through soil spaces.
Why overwatering can harm roots
Water fills soil air spaces and limits oxygen supply to roots.
Soil compaction effects
Reduces aeration and infiltration and increases runoff.
Root hairs
Root extensions that increase surface area for water and mineral absorption.
Concentration gradient
Difference in concentration that drives diffusion from higher to lower concentration.
Surface area and absorption
Greater surface area generally increases absorption rate.
Proton pump
Membrane protein that uses ATP to move H+ across a membrane.
Parasitic plant
Plant that obtains resources from a host plant.
Holoparasite
Parasitic plant that depends entirely on its host for nutrition.
Hemiparasite
Parasitic plant that obtains some resources from a host but can photosynthesize.
Mycoheterotroph
Plant that obtains nutrients entirely from fungi; typically lacks green coloration.
Insectivorous plant
Plant in nutrient-poor soil that obtains minerals, often nitrogen, from prey.
Mycorrhizal association
Mutualistic fungus-root association; fungus supplies nutrients and plant supplies carbohydrates.
Gas exchange in plants
Plants mainly take in CO2 and release O2 during photosynthesis; they also use O2 in respiration.
Why plants lack specialized gas-exchange organs
Each part can exchange gases; respiration is relatively slow; living cells are near the surface; air spaces aid diffusion.
Cuticle
Waxy layer that protects the plant and reduces water loss.
Palisade mesophyll
Tightly packed, column-shaped leaf cells rich in chloroplasts; major photosynthetic tissue.
Spongy mesophyll
Loosely arranged leaf cells with air spaces that facilitate gas exchange.
Stomata
Small openings, often on the lower leaf epidermis, that allow gas exchange and water-vapor loss.
Guard cells
Cells that control stomatal opening and closing.
Transpiration
Evaporation/loss of water vapor from plant surfaces, especially leaves.
CO2 in plants
Released by respiration but used in photosynthesis; usually diffuses from air into leaves.
O2 in plants
Released by photosynthesis and used in cellular respiration.
Guard-cell turgor
Water-filled guard cells become turgid and open stomata; loss of water makes them flaccid and closes stomata.
Stomatal opening sequence
Light activates ATP production and H+ pumps; H+ exits; K+ and Cl− enter; water potential decreases; water enters guard cells; stomata open.
Stomatal closing sequence
Abscisic acid signals ion release; solute conditions change; water leaves guard cells; cells lose turgor; stomata close.
Abscisic acid and stomata
ABA promotes stomatal closure during low light or low water availability and helps limit water loss.
Factors affecting stomata
Light, temperature, humidity, and CO2 concentration can affect stomatal number or activity.
Lenticels
Pores in stems or other tissues that allow gas exchange where cork layers block diffusion.
Homeostasis
Maintenance of relatively stable internal conditions despite external changes.
Regulator
Organism that maintains a relatively stable internal condition despite environmental variation.
Conformer
Organism whose internal condition changes with the external environment.
Trade-off in environmental tolerance
Coping with stressful conditions uses energy that could otherwise support other functions.
Plant hormone
Chemical signal that triggers a specific response in cells with the appropriate receptor.
Auxin
Hormone involved in phototropism, gravitropism, apical dominance, cell elongation, root growth, fruit development, and adventitious roots.
Phototropism
Plant growth response to light; auxin accumulates on the shaded side of a shoot and stimulates elongation there.
Apical dominance
Auxin from the shoot tip inhibits lateral bud growth; removing/pinching the tip can stimulate branching.
Cytokinin
Hormone that promotes cell division and growth, often in interaction with auxin; delays senescence.
Gibberellin
Hormone that promotes shoot elongation, leaf growth, seed and bud germination, and release from dormancy.
Abscisic acid (ABA)
Stress/dormancy hormone; inhibits growth, promotes seed/bud dormancy, and closes stomata during drought.
Ethylene
Volatile gaseous hormone that promotes fruit ripening, flower wilting, and leaf abscission.
Abscission
Shedding of a leaf, flower, or fruit from a plant.
Senescence
Aging process in plant tissues.
Apical meristem
Growth region at root and shoot tips that increases length.
Lateral meristem
Growth tissue that increases thickness or girth.
Intercalary meristem
Meristem at monocot leaf bases or nodes that enables continued growth after cutting.
Dermal tissue
Outer protective tissue covering the plant.
Vascular tissue
Tissue that transports water, minerals, and sugars; includes xylem and phloem.
Ground tissue
Tissue involved in photosynthesis, storage, and support.
Parenchyma
Common living ground tissue with thin walls; photosynthesis, storage, and wound repair.
Collenchyma
Living support cells with unevenly thickened walls, often beneath epidermis.
Sclerenchyma
Support tissue usually dead at maturity, with thick lignin-strengthened secondary walls.
Node
Point on a stem where leaves, aerial roots, or flowers attach.
Internode
Stem region between two nodes.
Primary growth
Increase in length due to apical meristem activity.
Secondary growth
Increase in thickness/girth due to lateral meristem activity.
Taproot system
One main deep root with smaller lateral roots; common in dicots.
Fibrous root system
Dense, shallow network of roots; common in monocots and helps reduce soil erosion.
Root cap
Protective covering at the root tip.
Root zone of cell division
Region near root tip where meristem cells actively divide.
Root zone of elongation
Region where newly formed cells increase in length.
Root zone of maturation
Region where cells differentiate into specialized types.
Endodermis
Layer separating root ground tissue from the stele.
Casparian strip
Waxy region in endodermal cell walls that helps prevent unwanted materials from entering root vascular tissue.
Lamina
Leaf blade; broad, flat part of a leaf.
Petiole
Stalk connecting a leaf blade to the stem.
Midrib
Main central vein of a leaf.
Leaf margin
Edge of the leaf blade.
Monocot leaf venation
Usually parallel veins.
Dicot leaf venation
Usually branched or net-like veins.
Trichomes
Small plant hairs that can help deter herbivores.