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primary aquatic orders
all members
immatures of ephemeroptera, odonata, plecoptera, trichoptera, megaloptera
secondary aquatic orders
some members
semi aquatic or specialized: neuropteran, orthopteran, hymenoptera
methods of getting O2 under water
breathe air with a tube
bring a bubble of air with you
diffusion
gills
respiratory pigments
breathe air with a tube
many insects have modified appendages that allow them to breathe air
eg: Nepidae
bring an air bubble with you
physical gill, as the oxygen gets depleted in the bubble, O2 from the water diffuses in and when its too depleted go to surface with a new bubble
diving beetles that hold a bubble of air under their elytra over their spiracles
dysticidae,hydrophilidae,halipidae
diving bugs that hold a bubble with airs on their ventral side
notonecidae, corixidae
fast flowing cold water means oxygenated water is constantly replenished around organisms
diffusion
gills work by
increasing surface area for diffusion
respiratory pigments allow
insects to store oxygen
eg hemoglobin
lotic habitat characteristics
flowing water
high O2 content
high habitat heterogeneity
external inputs such as leaf litter and wood
banks have tree-vegetated riparian zone
riffles
shallower,stony, fast-flowing sectionsp
pools
deeper areas
erosion
sediments taken from solid and carried away in water
those that live in strong currents are dorsoventrally flattened
often with laterally projecting legs
silk production
often live in benthos of streams between peebles and rocks
characteristic fauna of lotic habitats
lentic habitat characteristics
standing water
sizes vary from vernal pools to ponds and lakes
low O2 content
low habitat heterogeneity
limnetic zone
penetrated by ligth
littoral zone
near the shore where vegetation can grow
profundal zone
little to no light
insects adapted to water surface habitats
most within littoral zone
hydrofuge hairs
tarsal claws
detergent propulsion
divided eyes
insects adapted to pelagic zone
most in limentic zone
phantom midges, water boatman
insects adapted to profundal zones
chironomidae: non-biting midges
shredders
break up leaf litter and dead wood
collectors
filter-feeders on suspended particles
scrapers
graze algae off of solid surfaces
predators
feed on all other feeder types
river continuum concept
describes how stream ecosystems vary from headwaters to downstream and lastly to large ricers
upstream characteristics
highly shaded
narrow bed
low primary productivity
ecosystem respiration high
downstream characteristics
high light
wide bed
high primary productivity
lower ecosystem respiration
headwater characteristics
steep slopes
limited photosynthesis
leaf litter inputs
macro-invert diversity
springe, seeps-cold temps
transfer zone characteristics
some shading
channel widens and deepens
algae, submerged vegetation
water velocity increases
temperature, turbidity, and sediments increase
depositional zone characteristics
energy supplied by upstream DOM
diverse flood plain
omnivore fish species
drifting zooplankton and phyotplankton
integument
the outer covering of the living tissues of an insect
function of integument
water loss prevention
protection
support
regulates O2 flux
muscle attachment and movement
3 layers of integument
basement membrane
epidermis
cutile
epidermis
single cell layer of secretory tissue that produces the cuticle and part of basement membrane
regulates molting, type of cuticle secreted and placement of specialized cells
epidermis
inner sheet of connective tissue that separates the hemocoel from the integument and supports epidermal cells
basement membrane
cuticle
chemically complex layer that varies among and within insects
and consists of chains of chitin embedded in a protein matrix
tough,outer,acellular, secreted by cells of epidermis that is insoluble in water, dulites acids dilutes alkalis, alcohol, and all organic solvents
cuticle
3 layers of cuticle
endocuticle
exocutile
epicutile
procuticle
soft, chitin-rich cuticle, newly secreted by epidermis
exocuticle+endocuticle
exocuticle
scerletized cuticle
sceritization
process of hardening and darkening in exoskeleton via modification of the structure of both the proteins and chitin fibers
scleritization requires
amino acids tyrosine or phenylalanine
endocuticle
consists of unscelortized inner part of the procuticle that remains flexible
epicuticle
created by secretions from multiple sources deposited outside procuticle
what do dermal glands do to protect the wax layer of the epicuticle
secrete a cement atop the epicuticle
mix of hydrocarbons,alcohols, and fatty acids
wax
function of wax
prevent water loss
helps walk on water
pathogen resistance
insecticide resistance
camouflage
deter predation
communication
function of color production
protection from predation, solar/uv radiation
color production communication works by
attracting mates
thermoregulation
intra and inter individual recognition
2 mechanisms of color production
physical (structural colors)
pigmentary colorsp
pigmentary colors
results from absorption of visible light by chemical compounds
physical (structural colors)
result from light scattering, interferences, and diffraction
derived from protuberances on cuticle
physical colors
spines
mutlicellular, undifferentiated epidermal cells
setae
hairs- multicellular specialized cells
sensory of tactile environment
often modified into scales
acanthae
unicellular sclerotized protuberances that are hollow inside and some have pores under terminal plate for secretion
microtrichia
subcellular with several extension per cell, hairlike: solid cored of exocuticle covered by thin layer of epicuticle
4 main types of structural colors
spines
setae
acanthae
microtrichia
not metallic or iridescent, but vary enormously and are often derived from excretory products not involved in biochemical pathways
pigment colors
3 main tagmata of insects
head, abdomen, thorax
head
sensory perception , neural integration, and food gatheringth
thorax
for locomotion and machinery
abdomen
houses the visceral organs including components of the digestive, excretory and reproductive systems
insect tagmata evolution
segmented worms→arthropods→ insects
function of head
holds feeding appendages
hold important sense organs
contains and protects the brain
whats on the head
mandibles
specialized mouthparts
antennae
compound eyes
sometimes ocelli and other specialized features
prognathous
mouthparts directed forward and project anterior toward eyes
hypognathous
mouthparts directed downward relative to body, immediately ventral to head capsule
opisognathous
sucking beak is directed toward rear, beneath thorax
mandibulate
feeding appendages that are mostly complete, freely movable and not united into beak
parts of mouthparts
labrum
mandibles
maxillae
labium
labrum
upper lip
contains chemosensory sensilla
mandible
heavily sclerotized jaws that represent limbs that lost their distal segments
inner edge modified for biting
maxillae
2nd pair of jaws that are less massive and retain segmentation of appendages: sensory function
labium
lower lips
sensory function: palps of maxilla and labium function to taste and feel food
human importance of mouthparts
disease transmission and effectiveness of pesticides
4 main types of mouthparts
mandibulate, piercing-sucking, nonpiercing-sucking, mouthhooks
functions of mandibulate
hold,catch, manipulate and bite food
defense
courtship
construction of nests and shelters
mandibulate predators have
sharp apical teeth
mandibulate herbivores have
obtusely angled cutting teeth and basal grinding molars
piercing sucking mouthparts
feeding appendages where one or more are modified into a tubular organ for consuming liquid food
functions of piercing sucking mouthparts
pierce integument and suck fluids
nonpiercing-sucking mouthparts
feeding appendages where one or more modified into a tubular organ for consuming liquid food
functions on nonpiercing sucking mouthparts
to intake water and dissolved nutrients
suck fluids (nectar,honeydew, sweat)
mouthooks
larvae of diptera have highly reduced and invaginated head into thorax
move vertically
larvae with mouthooks feed on
variety of soft, semi liquid foods, including decaying matter and living plant and animal tissue
advantages of compound eyes
field of view:many insects can see in every direction
dual vision: see both dark, light, and color
disadvantages of compound eyes
resolution is lower
ommatidia
sensory cartridges in compound eyes
ocelli
simple eyes that are very sensitive to small changes in light
detect movement
stemmata
eyes in larval insects
type of simple eye
high light sensitivity, fast speed, and low resolution
3 type of antennae
scape
pedicel
flagellum
function of antennae
sensory
communication
movement detection
johnstons organs
mass of sensory cells that detects movement in antennae located within pedicel
3 segments of thorax
prothorax, mesothorax, metathorax
spiracles
openings for gas exchange
present laterally on mesa and meta thorax
6 main segments of legs
coxa
trochanter
femur
tibia
tarsus
pretarsus