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Types of pollinators
Honeybees, wildbees, bumblebees, butterflies, moths, bats, ants, beetles
How are pollinators important to wellbeing
Help our plants to reproduce (high yield crops), maintain healthy water and soil(plants), and air quality
What pollinators need
Collect nectar to produce food for offspring/larvals food plants
homes(soil, wood, hollow stems), food, water
How we can help pollinators
Create habitats for bees, avoiding insecticides and pesticides and growing native wildflowers
% flowering plants pollinated by animals
85
Insect pollination
34 billion in value pollinating crops, increase soybean yields 20 percent, blueberry over 200 percent, chocolate and pumpkins solely dependent on pollinators
Pollination
transfer of pollen from male to female parts in seed plants
self-pollination
same plant, easier for plant, greater probability of pollination
cross pollination
different plant, genetic recombination, gets brought to multiple plants, avoid inbreeding
Pollen vectors
wind, water, animals
Traits that let plants attract pollinators
Rewards: nectar, pollen, scented oil
Signals: colors, large size, shape, scent
Traits of effective animal pollinators
Surfaces/texture pollen can stick to, long tongue to access nectar, move from flower to next(speed/endurance/motivation), vibrate at certain frequency(buzz pollination), senses (sight/smell)
Color vision: Humans
3 types of cone cells
Perceived color depends on relative response of each type
visible light spectrum is 380-740nm(violet-red), shorter=UV, longer=infrared
Color Vision: Birds
Have 4 types of cone cells and can see UV through red lights
Color Vision: Insects
number of photoreceptor types varies between species, honeybees have 3 types from UV-orange, butterflies can have up to 15 types, less information about pollinating flies, around 10 types with 4 for color
Bees
White, UV, Yellow, Blue
Fresh, mild pleasant
Usually nectar
limited sticky pollen
shallow tubular landing platform flower shape
Beetles
Dull while or green
no odor/very fruity or bad smelling
sometimes nectar
ample pollen
bowl-like flower shape
Bird
Orange red white
no odor
ample, hidden nectar
modest pollen
large, tubular, strong perch
Wind
Dull green or brown
no odor
no necar
abundant, small, not sticky pollen,
small petals absent/reduced
Animal Pollination
Generally mutulistic, plant benefits(pollen carried further, could make less pollen(using less nitrogen) because more specific pollination
Animal benefits: food/nutrients from pollen and nectar
Plant costs: animal eats some pollen, plant sucess is tied to animal sucess, must produce nectar/flowers and attractive materials(cant use resources in other ways)
Animal costs: time and evergy going from one flower to another
Generalists vs specialists
-G: more widespread pollination, more options
-S: less competition, pollinators deliver pollen to another plant of same species and better reproduction
Ecology
how organisms interact with environment
environmental science is chem, geology, meteorology
Environmental studies: ethics, culture, clean energy, energy efficiency (aim to understand human impacts and mitigation)
Ecology falls within these, as they draw upon ecoloogy for info
Levels of biological organization
Individual
Population: A group of individuals of a single species that live in a particular area and interacting with each other
Community: association of pops of different species living in the same area at the same time
Ecosystem: a community of organisms plus their physical environment
ecological studies often include both biotic and abiotic components of natural systems
Landscapes: areas with substantial differences typically including multiple ecosystems
Biosphere: all living organisms on earth, plus the environments in which they live
Experiment
Scientists manipulate or change some variable to determine how it affects some outcome
Observational Studies
Scientists measure existing variation in nature to find out if theres a relationship between the two variables
Experimental Designs
Treatment: factor/variable you’re testing
Experimental group: factor you’re testing
Control group: doesnt have factor youre testing
Experimental unit: smallest unit to which a treatment is given
Confounding factors: variables other than the one you were testing that could affect the outcome
How to minimize confounding factors
Control as many as you can:
Replicate: apply each treatment to several experimental units
Randomize: assign treatments to experimental units randomly
Block: group similar experimental units together and apply one treatment to each experimental unit in the block
Case Study: Amphibian Limb Deformities
pacific tree frogs and salamanders with extra/missing limb, all had a parasite, implemented small beads near limb buds, beads caused similar deformities, collected eggs from ponds with and without parasites, pesticide combined with parasite had the most amount of deformities
Hypothesis
possible explanation for a phenomenon
Prediction
What you expect to see if your hypothesis is correct
Weather vs climation
-common: temp precip, humidity, wind, air, pressure, clouds
-differences: weather is short-term, climate is long term
climate: long term trands in temp, wind, and precip(daily, seasonal, and multi-year cycles - averages and variablility)
Climate affects
physiological function of organisms, resources available(water), abiotic processes such as rock weathering, which releases nutrients, rate of periodic disturbances like fires and mudslides
-climate determines the geographic distribution of organisms(often extremes more important than averages)
Differences in light energy drive temp, precip, wind patterns, andn ocean currents
Suns rays hit equator squarely, but hit edges further and angles
-equator gets much more sunlight than poles
Temp→global atmospheric circulation
solar radiation heats earths surface, warms air above it, warm air rises, expands, and cools, vapor condenses into clouds
Hadley cells
warm moist air rises in tropics, cools and drops moisture as rain. Cooler, drier air flows towards poles, this air descentds at about 30 degrees N and S, where deserts are located
Equator
O degrees, low pressure Tropics (30 degrees N and S high pressure)
Poles
Cold dense air subsides at poles, making polar cells. Polar cells and Hadley cells drive mid-latitude Ferrell cells

Air cells
Warm air rises, coolds and spreads out, sinks, precipitates near equator. Moise conditions in areas air rises, cold dense sir sinks.
Temp→global atmosphere circulation→precipitation
drives moisture/dryness
Temp and Precip→vegetation
Global atmospheric circulation → surface winds
coriolis effect converts winds moving to equator into east winds, and winds moving away into west winds
-east winds come FROM east, west, etc
Winds→Currents
when gulf stream reaches Arctic, water loses heat, becomes dense and saltier, and sinks
upwelling: brings nutrients from deep sediments to lighted surfacae waters wehre phytoplankton frow , food for zooplankton and fish
- prevailing wind blows parallel to coast, wind and coriolis effect creates surface current going away from coast
-deep cool water rises to replace the surface water
How seasonal climate variation caused by tilt of earth axis and its motion around the sun
because of earths axial tild - areas near equator usually have wet adn dry seasons, areas farther from equator have hot and cold seasons
How oceans, continents, and mountains afeect regional and local climate
water has high heat capacity - takes a lot of energy to change its temp
-maritime climate near coasts-low temp variation (duluth)
-continental climates-high temp variation (Day to day seasonal, year to year), seasonal variation in morris(avg low vs high), year to year (one jan to next jan)
Mountains - ways they affect climate
Elevation: higher=colder
aspect: sunnier side is hotter and drier - in N Hemisphere, south side is hotter and dried, N+E sides wooded, S+W sides dry with shrub
Wind direction: rain shadow on leeward, moving air gets moisture over oceans, is forced up over mountain, cools, and high elevation and falls as rain on side wind is coming from
rain shadow: dry side of mountain, after air already rained
Slide 43 mtn slide:
Temp patters explained by rockies and appalacions (cooler temp), less precip on rain shadow by rockies
MN climate
Normal annual precip: gradual increase precip going NW to SE, vertical differences, Normal mean temp: increase in temp going N to S with twin cities warmer on avg.
urban areas warmed by asphalt, brick without trees
warmer along N shore of Lake Superior than rest of nearby NE inland (pseudo-maritime)
NE MN cooler than NW (elevation)
—Warm humid summer/cold snowy winters: explained by earths tilt and 45 degrees from equator
—-Most precip in summer: moisture from gulf of mexico, warm air more moist than cold
— high variability day-to-day, seasonally, and year to year, cold as going N(curvature of earth, less solar in N)
—Moister as go SE, bc W MN falls in rockies rain shadow
Homeostasis
self-regulating process by which biological systems maintain stability while adjusting to changing external conditions
-self regulation includes negative feedback loops
Neg feedback loops
a change in one direction → change in the other direction negating initial change
-body temp increases, sweat to cool, temp decreases
Temp affects
enzyme stability(denatured protein), cell membrane fluidity, rates of biochemical reactions, freezing, water loss
How heat balance affects fuctioning of org
To survive changes in external temp, org must keep its internal temp constant(homeostasis) OR tolerate changes in internal temp
Warm blooded
Stable body temp = homeothermy
relies on internal heat production = endothermy
Cold blooded
Variable body temp = poikilothermy
relies on external heat production = ectothermy
Heat Balance
delta H = inputs-outputs
Oxidative metabolism
aerobic metabolism = C6H12O6 +6O2 → 6H2O + 6CO2 + energy
Plants can modify heat exchange
stomata closed (retain heat, not exposed), stomata open (cooler → evaporation)
Human thermal tolerance range
At temp outside optimal range (thermoneutral zone), must expend energy to maintain a stable body temp
Limits beyond which humans cannot maintain a stable body temp w/o external aids
upper limit for young, healthy people (111-118 F at 20 humidity, 87 F for 100% humidity)
Evaporative loss is huge for cooling humans, as humidity removes ability to cool (sweat)
Water Balance
Essential for life, H2O is medium in which biochemical reactions occur
water tied closely to salt balance
Water Homeostasis
Most organisms maintain relatively constant internal water and solute concentration
some desiccation-tolerant organisms can loser 80-90 percent water, go dormant and survive to rehydrate (like lichens, mosses, tardigrade)
Water balance: Delta W = inputs-outputs
Inputs: ingestion, oxidative metabolism
Outputs: segretion, evaporative loss
Either: osmotic exchange, driven by solute exchange → semipermeable membrane
most marine organisms have little osmotic exchange, move to freshwater: gain water - internal conc higher than external
marine mammals/bony fish put back in ocean, solute conc is lower in body, will lose water
Evaporative Losses
some loss inevitable for terrestrial organisms
necessary due to evaporative cooling, gas exchange, plants-transpiration moves mineral nutrients from roots to leaves
losses can be reduced by thick skin, waxy cuticle (tradeoffs)
individuals respond to stressful environments
immediate responses (physiology/behavior)
long term responses - acclimatization (physiology/morphology/changes in gene expression, no genetic change
Population response to stressful environment
Individuals can acclimatize; if genetic variation is present, pop can evolve by natural selection over several generations
called adaptation
change in allele frequency → genetic change
Adaptation is evolutionary process, acclimatization is NOT
Energy
radiant energy (vis light, infrared)
kinetic energy (motion, heat)
chemical energy - contained in chemical bonds (ATP, fats, etc), breaking bonds produces energy
Autotrophs
Produce own organic compounds that store energy
Photoautotrophs
energy from light
chemoautotrophs
energy from inorganic compounds (nitrifying bacteria)
Heterotrophs
Organic compounds and energy by eating others
detrivores
dead organic matter
Organic compounds
contain c-h bond
Photosynthesis net reaction
6H2O+6CO2+ light → 6O2 + C6H12O6 + sugar
glucose becomes nectar, starch, cellulose
What do organisms need to synthesize
Solar radiation, CO2, H2O, temp, nutrients
Tradeoffs affecting photosynthesis
Lead thickness: light vs room for cells
Stomata: water loss vs gas exchange(CO2 in, O2 out, CO2 reactant, O2 alternative reactant →photorespiration, no sugar produced, toxic products, costs energy, rubisco)
specialized forms of photosynthesis let plants take up CO2 at night or in separate cells from where sugars are made, but are less energy efficient
Biomes
Large-scale biological communities
shaped by physical environment, esp temp and precip
categorized by plant growth forms (trees, shrubs, grasses/evergreen vs deciduous/broadleaved vs needleleaved)
MN Biomes
Prairie Parkland - temp grassland(lower left)
Eastern Broadleaf Forest - temp deciduous forest (bottom right)
Laurentian Mixed Forest - boreal ( top right)
Tallgrass Aspen Parkland - (upper middle )
***locate dom vegetation of each one

Prairie Parkland temp and precip
Less precip and medium precip areas with more heat (evaporates water from soil)
Prairie Parkland plants adapted to temp and moisture
Dry/deep spreading roots, specialized photosynthesis, leathery/hairy leaves, deciduous leaves, succulent leaves/stems that store water
-wet soil - shallow roots, roots with air channels, prevents roots from getting O2 so roots stay shallow
Eastern broadleaf forest temp and moisture conditions
cold, deciduous leaves, sugar accumulation, starch storage
Laureation Mixed forest temp and moisture conditions
colder, needle like leaves, springy branches shed snow
Temperate grasslands
drought stress prevents tree growth
similar biomes will produce similar dominant species, even if across the world
Temperate deciduous forest
distinct seasons, temp variability, steady consistent precip, broadleaf deciduous tree
Temperate Shrublands and woodlands
Some months too low precip for plant growth, seasonal temp, woody evergreen shrub and scrub tree with thick leathery leaves
Temperate evergreen forests
Warm coastal region to cool continental maritime climates
varying precip
evergreen conifers and evergreen broadlead
Boreal Forest
Cold, short growing seasons and adequate precip, dominant plants are evergreen, needle-leaved tree
Deserts
High temp, low water availability, stem succulents help survive, can be hot/not hot, but all are dry
Tropical Rainforests
No real dry season
Tundra
Surface soil usually soggy
Temperate Grassland
Vast temporal shift, warm moist summer, cool dry winter
vast open, mainly grasses
moderate annual precip
Biome
Broad type of community, usually defined by dom vegetation
biome concept invented to classify terrestrial communities, can extend to aquatic environment, but doesn’t fit perfectly
ecosystem
species in particular place, plus abiotic environment they interact with
Lakes
Formed by glaciers or depressions moved by glaciers
Lake Superior
Largest freshwater lake by SA in world
oligotrophic (low and nutrient)
Severely P-limited (phosphorous), not much nutrients to support algae so not much fish (little food)
Oligotrophic Lake
Clear blue water, rocky bottom
eutrophic zone - enough light for photosynthetic organisms, 46-60m deep
aphotic zone - too dark for photosynthesis - superior avges 176 m deep
Mesotrophic lake
Medium nutrient levels
Eutrophic lake
High nutrient levels, not good, vegetation near shore, algae in water, euphotic zone shallow
Stratification = layering
Water most dense at 4 degrees C
warm surface water doesnt mix with cold deep water well, can cause low oxygen in deep water
Stratification and nutrient overload
algal growth on surface water, algal sink and decomposed in deep water, oxygen depleted in deep water, cant be replentished easily, gulf of mexico, southern MN lakes
Rivers
meanders and bends in rivers, fast vs slow flow
Wetlands around river
Slow streams down, allows plant growth and break down contaminents, store and slowly release water
Developed floodplains and humans want to live nearby river
levees can be eroded in hours, everything in floodplain can be flooded, upstream levees push water downstream
Bdote
where two waters came together