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Ecologists study…?
Ecologists study environmental relationships from individual organisms to factors that influence global processes.
What makes the transport of water through plants possible
Water cohesion
Physiological ecologists focus on…?
Physiological ecologists focus on the evolution of physiological and anatomical mechanisms that organisms adapt due to physical and chemical environmental challenges
Behavior ecologists focus on…?
Behavior ecologists focus on the evolution of behaviors that allow animals to survive and reproduce
Population ecology focuses on…?
Population ecology focuses on factors influencing population structures and processes
Population definition
Population: Group of interbreeding individuals of a single species in a defined area
Community definition
Community: Association of interacting species
Ecosystem definition
Ecosystem: Biological community together with its associated physical and chemical environment
Biosphere definition
Biosphere: The portions of the Earth that support life, including the land, waters, and atmosphere
What type of ecology is involved with the biosphere
Landscape ecology
What are the three types of research?
Observation, Experimentation, Modeling
What is observational research
Collection of data in unmanipulated settings
Give examples of observational research
Counting the number of birds or describing types of fungal spores
Work takes place either in situ, in the field, or using specimens collected in nature that are then observed in a laboratory.
Binoculars, drones with cameras, stable isotope analysis
What is experimentation research
Experimentation: Research that involves manipulation of variables of interest while holding others constant to test a hypothesis
Where does experimentation research occur?
Occurs in the field or controlled environments like a lab, garden, greenhouse, or outdoor enclosures
What is modeling research?
Modeling: Creation and analysis of representations of data or ideas to provide insight or make predictions
What are conceptual models?
Conceptual models: Systems in pictures or diagrams
What are quantitative models?
Quantitative models: Mathematical and may involve complex equations
What is a phenotype?
Observable characteristic
What is phenotypic plasticity?
Phenotypic Plasticity: Organisms change the expression of traits in response to the environment
What are ecotypes?
Ecotypes: Locally adapted and genetically distinctive populations within a species
Discuss the Clausen’s Common Garden Experiment
Individuals from two or more populations are transplanted and grown in the same environment. Cloning lowland, mid-elevation, and alpine plants and growing them in experimental gardens
Plant height differed significantly among the study sites. Different plants responded differently to the environments. Mid-elevation and alpine plants attained the greatest height in the mid-elevation garden. Lowland plants grew tallest in the lowland garden
What is the Hardy-Weinberg Principle for?
Hardy-Weinberg equilibrium: Helps identify evolutionary forces that can change gene frequencies in populations
What are the two equations for the hardy-weinberg pricnple?
p2 + 2pq + q2 = 1.0
p + q = 1.0
What are the conditions for the hardy-weinberg principle?
Conditions: Random mating, no mutations, large population size, no migration, and all genotypes have equal fitness
Fitness definition
Fitness: Genetic contribution of individuals to future generations
Natural selection definition
Differential survival and reproduction among phenotypes
Polygenic traits definition
Polygenic traits: Characteristics controlled by multiple genes
Traits may have two possible categories or vary along a gradient
Like skin color
Quantitative genetics definition
Quantitative genetics: Mathematical treatment of continuously varying traits and how they respond to natural selection
Stabilizing selection definition
Stabilizing selection: Acts against extreme phenotypes and can act to impede changes in populations
Stabilizing selection characteristics
Favors average phenotype
Bell-shaped curve
Average individuals have higher evolutionary fitness compared to extremes
Directional Selection defintion
Directional Selection: Favors extreme phenotype over other phenotypes
Disruptive Selection definition
Disruptive Selection: Favors two or more extreme phenotypes over the average phenotype in a population
Heritability definition
Heritability: The proportion of total phenotypic variation in a trait attributable to genetic variation. It determines the potential for evolutionary change in a trait
What is heritability symbolized as?
Usually symbolized as h2
Heritability equations:
h2 = VG / VP
h2 = VG / ( VG + VE )
h2 = VG / VP
h2 = VG / ( VG + VE )
What do the variables stand for?
h2 = Heritability
VG = Genetic variance
VP = Phenotypic variance
VE = Environmental effect
What does the heritability of a particular trait depend on?
Heritability of a particular trait depends on the relative sizes of genetic versus environmental variance
Describe Stabilizing selection for egg sizes
Eggs too small or too large lead to lower hatching and reduced lifetime production of offspring
Describe Directional selection for soapberry bugs
Feeds on seeds using slender beaks.
The beak length depends on the radius of the fruit. The larger the fruit, the larger the beaks
Describe Disruptive selection of finches
Favored smaller and larger beak sizes, not medium beak sizes
What can change gene frequencies in populations, especially small populations
Genetic drift
Define Genetic drift
Genetic Drift: Allele frequencies change over generations due to random chance instead of natural selection
Define founder effect
Founder effect: Subset of the original population becomes a new population
Define population bottleneck
Population bottlenecks: Large portions of a population are killed by a chance event unrelated to genotype, leaving behind reduced genetic diversity and/or different frequencies of alleles than existed before
What can population bottlenecks lead to?
Can lead to increased inbreeding
Populations with the highest level of inbreeding have the highest probabilities of extinction
Macroclimate definition
Macroclimate: Prevailing weather conditions in a region over a long period of time
Microclimate definition
Microclimate: Climatic variation on a scale of a few kilometers, meters, or even centimeters, usually measured over short periods of time
What are the six (kinda seven) things that affect microclimates?
Altitude: Temperatures are generally lower at higher elevations
Aspect: Topographic features such as hills and mountains due to shading some parts and exposing other parts to the sun.
Vegetation: Shades the landscape, creating microclimates, and plant litter also produces microclimates that decrease temperature
Color of the Ground: Changes the temperature
Presence of boulders and burrows: Decreases temperature
Aquatic temperatures:
Very stable temperature
High capacity to absorb heat
Large amount of heat absorbed by water as it evaporates
Heat energy that water gives up to the environment as it freezes
Riparian vegetation: Vegetation that grows along rivers and streams, influences the temperature in streams, decreasing temperature
What is the principle of allocation?
Principle of allocation: If an organism allocates energy to one function, such as growth or reproduction, it reduces the amount of energy available to other functions, such as defense
What did Bennett and Lenski test with bacterium fitness?
Bennett and Lenski tested bacterium fitness at 20C and 40C and found that adaptations occurred at 20C with decreased fitness compared to ancestors grown at 40C
What is acclimation?
Acclimation: Physiological adjustment to a change in a particular environmental factor, such as temperature or salinity
Equation for heat stored in the body of an organism
HS = Hm ± Hcd ± Hcv ± Hr - He
HS = Hm ± Hcd ± Hcv ± Hr - He
What do the variables stand for?
HS : The total heat stored in the body of an organism
Hm : Heat gained from metabolism
Metabolic heat is the energy released within an organism during cellular respiration
Hcd : Heat lost or gained by conduction
Conduction is the transfer of heat between objects in direct physical contact
Hcv : Heat lost or gained by convection
Convection is the process of heat flow between a solid body and a moving fluid
Hr : Heat lost or gained by radiation
Radiation is the transfer of heat through electromagnetic radiation,
He : Heat lost by evaporation
Define poikilotherms
Organisms whose body temperature varies directly with environmental temperatures
Define ectotherms
Ectotherms: Organisms that rely mainly on external sources of energy for regulating body temperature
Define endotherms
Endotherms: Organisms that rely mainly on internally derived metabolic heat energy
Define homeotherms
Homeotherms: Endotherms that use metabolic energy to maintain a relatively constant body temperature
Plant heat regulation eqation
HS = Hcd ± Hcv ± Hr
Metabolism is so small that it is negligible
To decrease overheating desert plants do what
Place their foliage far enough above the ground to avoid conductive heating.
Have small leaves, which give high conductive cooling
Have reflective surfaces to decrease radiative heat gain
To conserve heat, arctic/alpine plants do what
Have dark pigments and orient leaves and flowers perpendicular to sun rays, all of which increase radiative heat gain
Have cushion growth, which gains heat from warm substrate through conduction and reduces convective heat loss
What do Tropical Alpine Plants do to maintain temperature?
Retain dead leaves to insulate the stem, conserving heat and reducing convective heat losses
What do lizards do to maintain body temperature? What type of therm is it?
Ectothermic
Basks in the sun to warm its body or seeks shade to cool
What do grasshoppers do to maintain body temperature? What type of therm is it?
Ectothermic
Bask in the sun and can adjust capacity for radiative heating. Differing pigmentation between species of grasshoppers
What is the Thermal Neutral Zone? What type of therm animals use it?
Endothermic
The range of environmental temperature over which the metabolic rate of a homeothermic animal does not change
What happens when temperatures are too low and too high outside of the thermal neutral zone?
When temperatures are outside the zone:
Low Temperatures: Shivering, release hormones, etc
High Temperatures: heart rate and blood flow to skin increases which increases radiative heat loss
Why are aquatic animals able to be endothermic/homeothermic
Can be homeothermic in an aquatic environment because:
They are all air breathers and do not expose respiratory surface to the surrounding water
Well insulated with thick layers of fat. Parts not insulated rely on countercurrent heat exchangers
Describe how plants can be thermogenic
Use metabolic energy to heat their flowers
Can change metabolic rate depending on the temperature
How do many organisms survive extreme temperatures?
Entering a resting state
Define animal inactivity
Seeking shelter during the hottest or coldest times of the day
Describe animals reducing metabolic rate
Reducing Metabolic Rate: When food is scarce and night temperatures are low, an organism can enter torpor for short-term or hibernation/estivation for long-term
Torpor definition
Torpor: State of low metabolic rate and lowered body temperature
Hibernation definition
Hibernation: A dormant state, involving reduced metabolic rate, that occurs in some animals during the winter.
Estivation defintion
Estivation: A dormant state that some animals enter during the summer; involves a reduction of metabolic rate.
Discuss Hibernation in Tropical Species:
During the dry season of the tropics, which can last up to eight months, some organisms will hibernate.
What influences the movement of water between an organism and its environment
Concentration graidnets
Relative humidity defintion
Relative humidity: The percent of water content of air relative to a potential maximum
Relative humidity equation
RH = Water Vapor Density / Saturation water vapor density * 100%
Water vapor pressure definition
Water vapor pressure: The atmospheric pressure exerted by the water vapor in the air; increases as the water vapor in the air increases
Saturation water vapor pressure definition
Saturation water vapor pressure: The pressure exerted by the water vapor in air that is saturated with water vapor
Diffusion definition
Diffusion: Transport of material due to the random movement of particles; new movement is from areas of high concentration to low concentration
Osmosis definition
Osmosis: Diffusion of water down its concentration gradient
Aquatic organisms live in one of three circumstances. Name the three
Isomotic, Hypoosmotic, Hyperosmotic
Hypoosmotic definition
Hypoosmotic: Organism with body fluids with a higher concentration of water and lower solute concentration than the external environment
Isomotic defintion
Isomotic: Organism with body fluids with the same concentration of water and solutes as the external environment
Hyperosmotic defintion
Hyperosmotic: Organism with body fluids with a lower concentration of water and higher solute concentration than the external environment
Water potential definition
Water potential: The capacity of water to do work
How does changing solution concentration change water potential
Increasing solute concentration decreases water potential
Decreasing solute concentration increases water potential
What is the water potential of pure water?
0
What is the symbol of water potential?
Ψ (psi)
What is Ψsolutes ?
Ψsolutes is the reduction in water potential due to dissolved substances, which is a negative number
Matric forces defintion
Matric forces: A force resulting from water’s tendency to adhere to the walls of containers such as cell walls
Ψmatric
Cohesion defintion
Cohesion: A property of water whereby molecules stick together, making the transport of water through plants possible
Equation of water potential of plants?
Ψplant = Ψsolutes + Ψmatric + Ψpressure
How do Ψsoil and Ψmatric compare?
Ψsoil > Ψmatric
How do Ψplant and Ψsoil compare?
Ψplant < Ψsoil
How do terrestrial plants and animals regulate their internal water?
Terrestrial plants and animals regulate their internal water by balancing water acquisition against water loss
What is the animal water regulation equation?
Wia = Wd + Wf + Wa - We - Ws
Define variables of the animal water regulation equation
Wia = Wd + Wf + Wa - We - Ws
Wia : Internal water of an animal
Wd : Water taken by drinking
Wf : Water taken in with food
Wa : Water absorbed from the air
We : Water lost by evaporation
Ws : Water lost with secretion/excretion
What is the plant water regulation equation?
Wip = Wr + Wa - Wt - Ws