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_____ is sister to the rest of the Angiosperms
Amborella
ANA grade (paraphyletic)
Amborellales + Nymphaeales + Austrobaileyales
Vessels appear in the mrca of
Nymphaeales, Austrobaileyales, Magnoliids, Monocots and Eudicots

tracheid
vessel element
xylem
pits
perforation plate
true or false: amborella only have trachieds
true
Amborella and the reconstruction of an ancestral flower using extant taxa
Amborella’s morphology highly influences character reconstruction because of its position in the tree

Amborellales (family name? genus? 4 traits? part of ANA or Magnoliids)
Family: Amborellaceae
Genus: Amborella
Traits:
1. Tracheids
2. Undifferentiated perianth
3. Apocarpous gynoecium
-ANA
Amborella trichopoda
only member of Amborellaceae and Amborellales.

Nymphaeales (family, common name, traits, part of ANA or Magnoliids)
Family: Nymphaeaceae (water lilies)
Traits:
1. Aquatic herbs
2. Undifferentiated perianth
-ANA

order? family? common name? ANA or Magnoliids?
Order Nymphaeales
Family Nymphaeaceae
Common name: water lilies
ANA

order? family? common name? ANA or Magnoliids?
Order Nymphaeales
Family Nymphaeaceae
Common name: water lilies
ANA

Austrobaileyales (family, common name of family, ANA or Magnoliids)
Illiciaceae (star-anise family)
-ANA

order? family? common name of family? ANA or Magnoliids?
Order Austrobaileyales
Family Illiciaceae
Common name of family: star-anise
-ANA
Magnoliids are a clade sister to ___
the clade comprised by Monocots and Eudicots
* This position is still debated in the literature

Laurales (family, common name, traits, ANA or Magnoliids?)
Family Lauraceae
common names: Laurel, avocado, cinnamon family
Traits:
1. Trees and shrubs with aromatic oils
-Magnoliids
Who dispersed the avocado in nature?
Extinct (13 k years ago) megafauna including mammoths, horses, and giant sloths

Magnoliales (family, common name, traits, ANA or Magnoliids)
Family Magnoliaceae
common name: Magnolia
traits:
1. Trees or shrubs with solitary flowers
2. Undifferentiated perianth
3. Apocarpous gynoecium
-Magnoliids

Family Aristolochiaceae (order, traits, ANA or Magnoliids)
Order Piperales
traits:
1. Herbaceous, usually climbing
2. Enlarged petaloid calyx
-Magnoliids

Family Piperaceae (order, common name, traits, ANA or Magnoliids)
Order Piperales
common name: black pepper
Traits: 1. Ethereal oils
-Magnoliids
Basal Angiosperms: ANA + Magnoliids - Traits that frequently appear
1. Undifferentiated perianth
2. Apocarpous gynoecium
3. Ethereal oils
Ethereal oils / essential oils
• complex mixtures that may contain over 300 different compounds
• They consist of organic volatile compounds
Monocots apomorphies
1. Vascular cambium absent (no real wood)
2. Leaf venation parallel
3. One cotyledon
4. Embryo radicle (root) is reduced or ephemeral

Alismatales (family, common name, traits)
Family Araceae
common name: Arum family
traits:
• Spadix of numerous small flowers
• Subtending spathe
-monocots

(order? family? common name?)
Order Alismatales
Family Araceae
Common name: Arum family
-monocot

(order? family? common name?)
Order Alismatales
Family Araceae
Common name: Arum family
-monocot

(order? family? common name?)
Order Alismatales
Family Araceae
Common name: Arum family
-monocot

(order? family? common name?)
Order Alismatales
Family Araceae
Common name: Arum family
-monocot

Family Liliaceae (order, common name, genus, FF, trait)
Order Liliales
common name: lily family
Genus Lilium
FF: *, -3, 3-, 6, 3
Trait: Bulbous
-monocot

Family Melanthiaceae (order, common name, genus, FF)
Order Liliales
common name: Trillium family
Genus Trillium
FF: *, 3, 3, 6, 3 (fused and superior)
-monocot

Family Agavaceae (order, common name, traits)
Order Asparagales
common name: Agave family
trait: Xeromorphic, fibrous leaves
-monocot

order, family, common name
Order Asparagales
Family Agavaceae
Common name: Agave family
Order Asparagales, Family Agavaceae (Agave family) - Pollination process
• Adults emerge in spring and mate
• Pregnant females collect pollen and make into a ball
• Female deposit the ball of pollen into the stigma of flower
• Moth deposit its eggs in the ovary of the flower
• Eggs develop into larva, ovary develops into fruit
• Larva eat some of the seeds, then leaves the fruit and buries itself underground

Family Alliaceae (order, common name, traits)
Order Asparagales
common name: onion family
traits:
• Bulbous herbs
• Umbellate inflorescence
• Superior ovary
-monocot

Family Amaryllidaceae (order, traits)
Order Asparagales
traits:
• Bulbous herbs
• Umbellate inflorescence
• Inferior ovary
-monocot

Family Iridaceae (order, common name, traits)
Order Asparagales
common name: Iris family
traits:
• Unifacial leaves (equitant)
• Three stamens
• Inferior ovary
-monocot

Family Orchidaceae (order, common name, traits)
Order Asparagales
common name: Orchid family
traits:
• Labellum (inner tepal)
• Column, gynostemium, 1 fertile stamen fused with style and stigma
• Inferior ovary
-monocot
Column
1 fertile stamen fused with style and sigma
(filament + anther + style + stigma)


anther cap
viscidium
stigma
pollinia

pollinia
stipe
viscidium
Pollinarium
Pollinated stigma

Order Arecales (family, common name, traits)
Family Arecaceae / Palmae
common name: palm family
traits:
• Arborescent stem, monopodial
• Large plicate leaves
-monocot

order? family? common name?
Order Arecales
Family Arecaceae / Palmae
common name: palm family
-monocots