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Basal Angiosperm Families
ANA: Amborellaceae & Nymphaeaceae
Magnoliids: Magnoliaceae & Aristolochiaceae
Family Amborellaceae
Only has Tracheids & No Vessels
Spiraled Perianth
Dioecious: Unisexual Plants
Laminar Stamens
Apocarpic Pistils

Family Nymphaeaceae
Hollow Stems with Only Tracheids & No Vessels
Air Cavities in Tissue
Floating & Submerged Leaves
Showy Flowers with Many Parts
Laminar Stamens
Superior Syncarpic Pistil
Floral Formula: CA4-♾ CO♾ A♾ G(♾)
Genera: Nymphaea
Family Magnoliaceae
Large & Pinnate Veined Stipulate Leaves
Large & Solitary Flowers with Spiraled Perianth & Many Parts
Laminar Stamen
Apocarpic Pistils
Produce an Aggregate of Follicles
P♾ A♾ G♾
Genera: Magnolia
Family Aristolochiaceae
Climbing or Rhizomatous Herbs
Cordate Leaves
Flower with Petaloid Calyx
Hypanthium
Absent or Heavily Reduced Petals
Genera: Asarum
Genus Magnolia (Magnolias)
Woody Tree or Shrubs
Large Showy Flowers with Many Spirally Arranged Tepals & Stamens
Superior & Apocarpic Pistil with Many Carpels on Elongated Receptacle
Simple & Alternate Leaves with Pinnate Venation
Aggregate of Follicles as Fruit


Floral Formula of Magnolia
P∞ A∞ G∞
Genus Nymphaea (White Water Lily)
Aquatic Herb
Simple Floating Leaves with Notched Cordate Base & Palmate Venation
Large Showy Flowers with Many Tepals & Laminar Stamens
Semi-Inferior & Syncarpic Pistil with Many Carpels
Spongy Berry-Like Fruit
Mucilaginous Tissues & Strong Floral Scent


Floral Formula of Nymphaea
P∞ A∞ -G-(∞)
Genus Asarum (Wild Ginger)
Low Rhizomatous Woodland Herb
Simple & Basal Cordate Leaves with Pinnate Venation
Solitary & Mauve-Colored Flower with Petaloid Calyx & Absent Corolla
Hypanthium Present
Inferior & Syncarpic Pistil with 6 Carpel
Capsule Fruit


Floral Formula of Asarum
CA(3) CO0 A12 -G-(6)
Family Ranunculaceae
Perennial Herbs
Usually Alternate Leaves with Palmate Venation
Many Stamens & Carpels
Genera: Caltha, Clematis
Genus Caltha (Marsh Marigold)
Wetland Herb
Simple & Basal Cordate Leaves with Palmate Venation
Bright Yellow Flowers with Petaloid Calyx & Absent Corolla
Superior & Apocarpic Pistil with Many Carpels
Aggregate of Follicles Fruit


Floral Formula of Caltha
CA5 CO0 A∞ G∞
Genus Clematis (Clematis)
Vine or Climbing Herb
Opposite & Usually Compound Leaves
Showy Flowers with Petaloid Calyx & Absent Corolla
Superior & Apocarpic Pistil with Many Carpel & Long Feathery Styles
Feathery Aggregate of Achenes


Floral Formula of Clematis
CA4-8 CO0 A∞ G∞
Family Berberidaceae
Small Shrubs or Herbs
3-Merous Flowers
Marginal Placentation
Genera: Beberis, Podophyllum
Genus Berberis (Barberries)
Shrub with Spines
Simple & Alternate Leaves with Pinnate Venation & Often Clustered on Short Shoots
Yellow or White 3-Merous Flowers
Superior & Unicarpellate Pistil
Red Berries


Floral Formula of Berberis
CA3+3 CO3+3 A3+3 G1
Genus Podophyllum (Mayapple)
Woodland Herb
One or Two Simple & Peltate Palmately Lobed Leaves
Single 3-Merous Flower hanging Beneath Leaves
Superior & Unicarpellate Pistil
Berry Fruit


Floral Formula of Podophyllum
CA6 CO6/9 A6/9 G1
Family Papaveraceae
Herbs with Showy Flowers
Alternate & Compound Leaves
Deciduous Sepals & Crumpled Petals
Syncarpic Pistil
Capsule Fruits
Genera: Sanguinaria, Dicentra
Genus Sanguinaria (Bloodroot)
Herb with Orange-Red Latex from Rhizomes
Basal & Palmately Lobed Leaves
Two Deciduous Sepals
White & Actinomorphic Flowers with Many Stamen
Superior & Syncarpic Pistil with 2 Carpels
Capsule Fruit


Floral Formula of Sanguinaria
CA2 CO8-12 A∞ G(2)
Genus Dicentra (Dutchman’s Breeches)
Herb
Alternate & Compound Leaves that are Finely Divided
Two Deciduous Sepals
Strongly Zygomorphic & Somewhat Heart-Shaped Hanging Flowers with Spurred Petals
Superior & Syncarpic Pistil with 2 Carpels
Capsule Fruit


Floral Formula of Dicentra
CA2 CO4 A6 G(2)
Family Nelumbonaceae
Aquatic Herb
Peltate Emergent Leaves
Solitary Flower with Filamentous Stamens
Apocarpic Pistil with Many Carpel
One Carpellate Fruits in a Head
Genera: Nelumbo
Genus Nelumbo (Lotus-Lily)
Aquatic Herb
Simple & Peltate Leaves with Radial Venation that rise Above Water
Showy Flower with Many Tepals & Filamentous Stamens
Superior & Apocarpic Pistil with Many Carpels
Nut-Like Fruits embedded in Enlarged & Showerhead-Like Receptacle


Floral Formula of Nelumbo
P∞ A∞ G∞
Caryophyllid Characteristics
Betalain Pigments
Centrospermae: Free-Central or Basal Placentation
Dichasial Cyme Inflorescences Common
Adapted to Stressful Environments
Deserts, Saline Areas, Nutrient-Poor Bogs, Alpine Habitats
Family Caryophyllaceae
Herbs with Swollen Nodes & Opposite Leaves
Showy 5-Merous Flowers in Dichasial Inflorescences
Superior & Syncarpic Pistil with Free-Central Placentation
Capsule Fruit
Genera: Saponaria, Silene
Family Amaranthaceae
Herbs
Often Weeds & Halophytes
Tiny & Congested Flowers with Absent Corolla
Superior & Syncarpic Pistil
Utricle or Achene Fruit with Persistent Calyx
Genera: Amaranthus, Chenopodium
Family Cactaceae
Stem-Succulents
Spines/Glochids at Areoles
Absent or Reduced Leaves
Showy Flowers with Many Tepals & Stamens in Hypanthium
Inferior & Syncarpic Pistil
Berry Fruit
Genera: Opuntia
Family Phytolaccaceae
Shrubs
Small Flowers with Petaloid Sepals & Absent Corolla in Racemes
Superior & Semi-Syncarpic
Dark-Colored & Poisonous Berries
Genera: Phytolacca
Family Portulacaceae
Succulent Herbs with Fleshy Leaves
Bright Flowers with 2 Sepals & 5 Petals
Superior & Syncarpic Pistil
Pyxis Capsule Fruit
Genera: Portulaca
Family Montiaceae
Small & Fleshy Herbs with Basal Leaves
Delicate Flowers with 2 Sepals & 5 Petals
Superior & Syncarpic Pistil
Capsule Fruit
Genera: Claytonia
Family Polygonaceae
Herbs with Swollen Nodes & Ocrea
Alternate Leaves
Small 3-Merous Flowers with Tepals
Superior & Syncarpic Pistil
Three-Sided Achene
Genera: Persicaria
Genus Amaranthus (Amaranth & Pigweed)
Weedy Herbs
Alternate & Simple Leaves with Pinnate Venation
Tiny Green Flowers with Absent Corolla on Congested Spike Inflorescences
Superior & Syncarpic Pistil with 2 or 3 Carpels
Utricle Fruit


Floral Formula of Amaranthus
CA5 CO0 A5 G(2-3)

Genus Chenopodium (Lamb’s Quarter)
Tall Herb
Alternate & Simple Leaves with Goosefoot-Shape & Mealy White Coating
Tiny Green Flowers with Absent Corolla in Inflorescence
Superior & Syncarpic Pistil with 2 Carpels
Utricle Fruit

Floral Formula of Chenopodium
CA5 CO0 A5 G(2)
Genus Opuntia (Prickly Pears)
Stem Succulents of Flattened Cladodes
Leaves Absent
Tiny Glochid Spines in Tufts on Areoles
Large & Epigynous Flowers with Many Tepals & Stamens in Hypanthium
Inferior & Syncarpic Pistil with 4 Carpels
Berry Fruit


Floral Formula of Opuntia
P∞ A∞ -G-(4)
Genus Phytolacca (Pokeweed)
Shrub with Red-Purple Stems
Alternate & Simple Leaves
Small Pinkish-White Flowers with Petaloid Calyx & Absent Corolla in Long Racemes
Superior & Semi-Syncarpic Pistil with 5 to 10 Carpels
Dark-Colored & Juicy Berries


Floral Formula of Phytolacca
CA5 CO0 A10 G(5-10)
Genus Saponaria (Soapwort)
Somewhat Tall Herb with Swollen Nodes
Opposite & Simple Leaves
Showy 5-Merous Flowers with Tubular Fused Calyx & Blade-Claw Petals
Superior & Syncarpic Pistil with 2 Carpels
Capsule Fruit


Floral Formula of Saponaria
CA(5) CO5 A10 G(2)
Genus Silene (Campion)
Somewhat Tall Herb
Opposite & Simple Leaves
Showy 5-Merous Flowers with Inflated Fused Calyx & Deeply Notched Corolla
Superior & Syncarpic Pistil with 3 Carpels
Capsule Fruit


Floral Formula of Silene
CA(5) CO5 A10 G(3)
Genus Portulaca (Purslane)
Spreading/Trailing Succulent Herb
Alternate or Opposite Fleshy Leaves
Bright & Colorful Flowers with 2 Sepals & 5 Petals
Superior & Syncarpic Pistil with 3 Carpels
Circumscissile Pyxis Capsule Fruit


Floral Formula of Portulaca
CA2 CO5 A5-∞ G(3)
Genus Claytonia (Spring Beauty)
Ephemeral Herb
Two Opposite & Basal Leaves
White-Pink Flowers with 2 Sepals & 5 Petals
Superior & Syncarpic Pistil with 3 Carpels
Capsule Fruit


Floral Formula of Claytonia
CA2 CO5 A5 G(3)
Genus Persicaria (Smartweed)
Stemmy Herb with Swollen Nodes & Ocrea Present
Alternate & Simple Leaves
Small & Pink-Colored 3-Merous Flowers in Dense Spike Inflorescences
Superior & Syncarpic Pistil with 3 Carpels
Three-Sided Achene Fruit


Floral Formula of Persicaria
P3+3 A6-8 G(3)
Rosid Characteristics
Usually Separate Petals
Enormous & Diverse Clade
Usually 4 or 5-Merous
Family Oxalidaceae
Low Herbs
Compound Trifoliate Leaves
5-Merous Flowers
Often Tristylous
Explosive Capsule Fruit
Genera: Oxalis
Genus Oxalis (Wood Sorrel)
Low Growing Herb
Alternate & Trifoliate Leaves with Pinnate Venation & Ability to Fold
Yellow, Pink, White, or Purple 5-Merous Flowers
Superior & Syncarpic Pistil with 5 Carpels
Tristyly: 2 Sets of Anthers & 1 Set of Styles at Different Levels
5-Locular Capsule Fruit with Explosive Seed Dispersal


Floral Formula of Oxalis
CA5 CO5 A5+5 G(5)
Family Violaceae
Low Herbs
Bilateral Flowers with Spurred Petals
Syncarpic Pistil with Parietal Placentation
Genera: Viola
Genus Viola (Wild Violet)
Small Herb
Simple & Usually Basal Cordate Leaves with Pinnate Venation
Purple, White, or Yellow 5-Merous & Zygomorphic Flowers with Distinct Nectar Spur on Lower Petal
Superior & Syncarpic Pistil with 3 Carpels & Parietal Placentation
5 Stamens form Cone around Ovary
Three-Valve Capsule Fruit


Floral Formula of Viola
CA5 CO5 A5 G(3)
Family Malvaceae
Herb with Stellate Hairs
Palmately Veined Leaves
Somewhat Showy Flower with Valvate Sepals
Monadelphous Stamens fused into Staminal Tube
Genera: Malva
Genus Malva (Mallow)
Herb
Alternate & Simple Leaves with Round to Cordate Shape & Palmate Venation
Star-Shaped Stellate Hairs
Showy 5-Merous Flowers with Fused Valvate Calyx
Edges of Sepal Meet without Overlapping
Monadelphous Stamens fused into Tube around Style
Superior & Syncarpic Pistil with 5 to Many Carpels
Schizocarp Fruit with Mericarp Wedges


Floral Formula of Malva
CA(5) CO5 A(∞) G(5-∞)
Family Onagraceae
Somewhat Tall Herbs
Epigynous & Usually 4-Merous Flowers
Hypanthium
Inferior & Syncarpic Pistil
Genera: Oenothera
Genus Oenothera (Evening Primrose)
Somewhat Tall Herbs
Alternate & Simple Leaves with Pinnate Venation
Epigynous & 4-Merous Flowers with Distinct Floral Tube & Large Yellow Corolla in Hypanthium
Inferior & Syncarpic Pistil with 4 Carpels
Capsule Fruit


Floral Formula of Oenothera
CA4 CO4 A4,8 -G-(4)
Family Grossulariaceae
Woody Shrubs
Palmately Lobed Leaves
Sometimes have Spines
Epigynous Flowers with Well-Developed Hypanthium & Inferior Ovary
Berry Fruits
Genera: Ribes
Genus Ribes (Gooseberries & Currants)
Shrub
Alternate & Simple Leaves with Palmate Lobbing & Venation
Spines Sometimes Present
Epigynous & 5-Merous Flower with Well-Developed Hypanthium
Inferior & Syncarpic Pistil with 2 Carpels
Berry Fruit with Dried Floral Parts at Tip


Floral Formula of Ribes
CA5 CO5 A5 G(2)
Family Fabaceae
Nitrogen-Fixing Herbs
Trifoliate or Compound Leaves with Stipules
Pea Flower with Banner, Wings, & Keel Petals
Unicarpellate Pistil with Marginal Placentation
Legume Fruit
Genera: Baptisia
Genus Baptisia (Wild Indigo)
Perennial Herb with Blue-Green Foliage
Alternate & Trifoliate Compound Leaves with Pinnate Venation
Pea Flower with Banner, 2 Wing, and 2 Fused Keel Petals
White, Yellow, Cream, or Blue
Non-Fused Stamens unlike Most Papilionoid Legumes
Superior & Unicarpellate Pistil
Inflated Legume Fruit


Floral Formula of Baptisia
CA(5) CO5 A10 G1
Family Urticaceae
Herbs with Stinging Hairs
Opposite Leaves
Reduced Unisexual & Wind-Pollinated Flowers
Unicarpellate Pistil
Genera: Urtica
Genus Urtica (Nettle)
Herb with Stinging Hairs
Opposite & Simple Leaves with Pinnate Venation & Serrate Margins
Unisexual Flowers with Absent Corolla in Inflorescence
Superior & Unicarpellate Pistil
Achene Fruit


Floral Formula of Urtica
CA4 CO0 A4 OR CA4 CO0 G1
Family Fagaceae
Trees & Shrubs
Large & Pinnately Lobed Leaves
Wind-Pollinated & Unisexual Flowers in Catkins or Cupule
Acorn Nut Fruits
Genera: Quercus
Genus Quercus (Oak)
Woody Tree
Alternate & Simple Lobed Leaves with Pinnate Venation
Unisexual Flowers with Absent Corolla
Staminate/Male Flowers in Dangling Catkins
Carpellate/Female Flowers in Cupule of Fused Bracts
Inferior & Syncarpic Pistil with 3 Carpels
Acorn Nut in Cupule


Floral Formula of Quercus
CA4-7 CO0 A4-12 OR CA6 CO0 -G-(3)