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Taxonomic (Morphological)
based primarily on distinct morphological differences
Biological Species Concept
based on infertility (cross ability) among individuals. Yet this doesn't work for asexual species
Ernest Mayer
: responsible for the founder effect, formally defined the biological species concept as "a group of interbreeding natural populations that are reproductively isolated from such groups
Allopatric
mode of speciation, get divergents but then they come back together
Sympatric
mode of speciation within a population, you start to get genetic divergence, get gene flow between species and ancestors which stops divergence. This is quite rare in animals, occurs under specific circumstances
Premating
preventing zygote formation, Geographical, ecological, Temporal, behavioral, Mechanical, prevention of gamete fusion
Douglas Shemske
was particularly interested in these flowers with different than its typical, found premating isolation in species
Post-mating
: preventing the proper functioning of zygotes once they are formed. Inviability, sterility, or abnormal developments of hybrid, F2 breakdown
ex) mule
Hybridization
The exchange of genes between species as a result of occasional inter-specific mating. Varies among different groups of organisms, common plants, fish but rare in mammals, can result in complex patterns of variation
Polyploidy
: an organism, tissue or cell with more than 2 complete sets of homologous chromosomes. Polypoid conditions range from autopolyploidy (AAAA) to allopolyploidy (AABB
Allopolyploidy
(more common) arises occasionally from hybridization between species
Linnaeus
father of taxonomy, binomial nomenclature, hierarchical system: kingdom phyla class orders families genera species
Phenetics
classifying species based solely on overall resemblance. Now largely dead
Cladistics
classifying species on the basis of their phylogenetic relationships
Hennig
invented cladistics and birth of phylogenetic traits which could not provide a depiction of the evolutionary relationships but was a hypothesis.
Monophyletic Group
: single ancestor gave rise to all species in the taxon and no species in any other taxon
clinal pattern of variation
A gradual change in an inherited characteristic across the geographic range of a species, usually correlated with an environmental transition such as altitude, temperature, or moisture.
Non-monophyletic Group
taxon whose members are derived from 2 or more ancestral forms not common to all members
Homology
similarity of traits due to shared ancestry
Homoplasy
similarity of traits because of convergent evolution
Convergent evolution: the evolution of structures that resemble one another and perform similar functional roles due to the shared ecology of unrelated organisms.
W & D Maddison
DNA sequencing is enabling rapid construction of the tree of life. Won apple scholarship and made program and now developed tree of life
Biological invasions
Successful establishment of a species in a region not previously occupied followed by a rapid range expansion. Some are harmful/ some are not
Lome Wolfe
enemy release hypothesis: was interested in possibility that alien species release their enemies when they leave native land. If you go in native range, the level of pest and disease pressure was 17% higher in native range vs introduced, involved both specialist and generalists
Common Attributes of Successful Invasive Species
• Rapid development of reproduction
• Uniparental reproduction (cloning or selfing)
• High reproductive output
• Well-developed dispersal mechanisms
• Broad ecological tolerance
- high phenotypic plasticity
Controls of invasive species
• ¬mechanical (hand/machines)
• Chemical (herbicides)
• Ecological (burning/flooding)
• Biological control
Biological Control
Planned introduction of natural enemies (predators, parasites, pathogens) to control unwanted populations of invaded alien range
Art Weis EBB
studied field mustard seed in southern California in both wet year and dry year. Wet year had many flowers, dry year had barely any, huge year-year fluctuations
Directional selection on flower time
Evidence for directional selection for early flowering in both populations in 2002
Earlier flowering plants set more seed than later flowering plants
Biodiversity
the number and kinds of living organisms in a given area
Background extinction
turnover of species at a low rate, a natural feature of eco-systems; estimated at approx. 1 species/year
2 elements of biodiversity
1. Interspecific variation or species diversity
2. Intraspecific variation or genetic diversity
E.O Wilson
in principle we could fix environmental problems but can't fix the loss of biodiversity. Development is like Spielberg style creation, we barely know anything of it
Mass Extinctions
very large numbers of extinction due to natural catastrophe
Anthropogenic extinction
- caused by humans now estimated 4-6000 species a year (probably significant underestimate)
Minimal Viable Population Size (MVP)
The number of individuals necessary for a species to maintain or increase its number in a region. Smallest population of a species that can sustain itself in face of environmental variation.
SLOSS debate
Debate in ecology and conservation biology during 1970's as to whether a single large or several small (SLOSS) reserves were superior means of conserving biodiversity in fragmented habitat
A generalist species
is able to thrive in a wide variety of environmental conditions and can make use of a variety of different resources (for example, a heterotroph with a varied diet).
A specialist species
can thrive only in a narrow range of environmental conditions or has a limited diet
non-adaptive
lacking in adaptive significance
maladaptive
they reduce fitness, likely in changing environements
tradeoff
ex of Hare with large ears in cold environment, smaller ears=better heat regulation but you compromise hearing activity for predation
Principle of allocation
resources invested in one function unavailable to invest in other functions. The strong relationship between ear size has largely unconstrained by other factors
reproductive-survival tradeoffs
investing resources in offspring means they can't be used in maintaining the body
geographic range
the areas of the planet where the species many be found, usually depicted as range maps
biome level variation
places with more precipitation develop vegetation that is taller, and usually more species rich and productive than places that are drier
soil
formed by the action of living organisms and geophysical processes of some mineral substance that we call "parent material"
parent material
a geologic subject rather then ecological often bedrock, but could be sand deposited in an area by wind and water. becomes greatly altered and added to by biological processes and chemical actions
xeric
very dry soils
hydric
water saturated soil
mesic
intermediate soil
leaching
if rainfall is heavy, dissolved nutrients are carried far down deep aquifers in the soil, below reach of plants
humus
organic matter so decomposed source no longer recognizable
loams
best soils, contain some sand and some clay and plenty of organic matter. The organic matter helps to stay mesic and keep nutrients available
Posoilized soils
cold northern regions, tan, sandy acid and nitrogen depleted
Lateritic soils
lowland tropical, ancient, red clay rich, heavy in iron and aluminum, leached of good nutrients, can support impressive forests but bad for farming
Net primary productivity
refers to the amount of new biomass produced annually through plant growth
dog hair forests
when many seeds start growing at once, from a disturbance, competition for light increases and trees stunted by crowding need to be manually thinned out
terpen resins
are tress principal defense against specialized insects and herbivores
grazing
eating leaves
browsing
eating woody parts
flag trees
branches grow outward only by leeward side of trunk where some buds can escape damage
cushion plants
form dense hemispherical dome of leaves that allows wind to flow over the plant in a smooth flow
ecological speciation
when 2 populations of the same species get different local adaptations because of the different environments they're in, with different selection pressures. This leads to the 2 populations diverging and evolving until they become reproductively isolated and then 2 different species
main problem of Alpine tundra
alpine organisms need to adapt to rapid oscillation of extreme hot and cold and expose organism to extreme UV radiation
evotranspiration
total amount of water that evaporates from terrestrial surfaces or is transpired from plant leaves
potential evotranspiration
that amount of water that could be lost to the atmosphere if that amount of water were available in the system, PE is the measure of the drying up power of the environment
Intertropical convergence zone
aka the doldrums, area encircling earth near the solar equator where NE and SE trade winds converge. the 2 hadley cells meet and the air rises, because the air is so hot and buoyant, then it begins to rise and cool and condense and precipitate (countries in it at all times always get rain aka rain forests)
ecological convergance
plants adapted drip tip to prevent establishment of epiphyll crust, good ex of it
Janzen-Connell hypothesis
supposes that lack of winter in tropical areas means that plants never gain any respite from the attacks of herbivores insects and other enemies such as fungi
Tropical rainforest
is pollinated by animals versus deciduous and boreal is wind
Range shifts
climatically induced range shifts by resident species are usually slow. Dramatic range shifts can occur rapidly when a new species is introduced to region which can become invasive
Warbler- Kirtlands
a small insect eating bird that nests ONLY in forests pf jack pine that have grown after fires that occurred between 5 and 20 years before. This bird is a habitat specialist and very rare, cognitively limited, maladaptive
tropic of cancer
latitudinal lines around the earth at 23.5N
tropic of capricorn
latitudinal lines around the earth at 23.5S
solar equator
line of latitude closest to the sun, oscillates between the 2 tropics lines making 1 cycle per year
atmospheric circulation
solar heating of the bottom of the atmosphere sets up atmospheric circulation patterns
doldrums
windless equatorial area 0-60 going upward or 30 going down
horse latitudes
windless area 30N and 30S
Roaring 40's
too much wind, at 45S
jet streams
concentrated and narrow westerlies and wander in irregular fashion
oceanic circulation
massive packets of H20 produce circulation patterns called currents
Continental climate
large seasonal temperature differences, with warm to hot (and often humid) summers and cold (sometimes severely cold) winters. Usually in the middle of a continent, no body of water to moderate temperature
Maritime climate
typical of the west coasts at the middle latitudes of most continents, and generally features warm (but not hot) summers and cool (but not. cold) winters, with a relatively narrow annual temperature range. Oceans provide thermal inertia which moderates the temperature, great lakes also induce this
orogenic precipitation
mountain generated precipitation
rain shadow
humid air gets pushed by prevailing westerlies to the East from the Ocean. It reaches the mountains and rises. Upon rising it gets cold, condenses and rains/ snows on the mountain. Therefore once air hits the other side of the mountain its dry. Thus you get a large patch east of the mountain where its dry and not a lot of vegetation
limiting factors
factors most important in determining whether a species cannot persist in an area
environmental gradient
the simplest sort of gradient would occur along s line drawn across a real habitat ex) walking from shore to higher ground
thermoconformers
whose bodt temps are more closely track ambient temperatures (fish, amphibians, reptiles)
conduction
direct transfer of heat between 2 bodies that are in contact
convection
heat transfer facilitated by a moving fluid usually air or water
radiation
heat transfer by electromagnetic radiation
redistribution
circulatory system redistributes heat among body parts especially core appendages
SA/V ratio
the way heat is lost only varies with SA not volume
small things have higher SA/V ratio, volume determines how much heat something can hold
Bergmann's rule
homeotherms tend to be larger at higher altitudes (cod) ex) bear, polar bears have big body to hold heat but little SA as more surface area the more heat you lose to environment
Allen's rule
homeotherms tend to have smaller appendages at higher altitudes (colder), they try to have minimum SA/V ratio, like a sphere so they can keep warm. but in hot climates max SA/V ratio try to be flat so they can get rid of heat
countercurrent circulation to limbs
arteries and veins should be appressed in appendages to conserve heat; separated in appendages designed to shed heat.
maintains gradient so heat is always flowing from outgoing to incoming blood
Skinny weasel in cold climate: tradeoff
being long and thin makes weasels subject to thermal stresses, but they need to be thin so they can go into the burrows of their prey, a compromise for food