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adaptation
heritable characteristic that enables organism to thrive in environment
biological evolution
heritable change in a population over time
lamarck
environmental changes generate organismal needs and structures become modified through use and are passed down
natural selection
differential reproduction and survival of individuals that results in elimination of maladaptive traits from a pop
variation among individuals within a pop in some heritable characteristics
variation results in differences among individuals in their fitness
fitness
genetic contribution by an individual’s descendants to future generations; proportionate contribution of an individual to future generations
phenotypic plasticity
changing gene expression in response to changing environment conditions
-ability of a genotype to produce more than one phenotype when exposed to different environments
reaction norm
range of phenotypes produced by single genotype
developmental plasticity
a type of phenotypic plasticity —> not reversible
acclimation
phenotypic plasticity in response to current envi conditions that is eversible
ex: seasonal changes in temperature tolerance in fish
phenotypic evolution
change in mean or variance of a phenotypic trait as a result of change in allele frequencies
directional selection
when mean value of a trait is shifted toward on extreme
stabilizing selection
natural selection favors individuals near population mean value of a trait, at the expense of those at either extreme
disruptive selection
natural selection favors individuals at each extreme at the expense of mean
cline
measurable, gradual change over a geographic region in some phenotypic character or a gradient in genotypic frequency
ecotype
population adapted to its unique local environmental conditions
abrupt change from other pops of same species
geographic isolate
prevented by some intrinsic barrier from interbreeding with other sub pops
allopatric isolation
complete geographic isolation
adaptive radiation
evolution from common ancestor of divergent forms adapted to distinct ways of life
oxygenic photosynthesis
release of oxygen
light and dark reactions
light reactions
require sunlight as energy source —> split water
chlorophyll as light absorbing pigment
split water
nadp+ —> nadph
adp —> atp
dark reactions
no light required
carboxylation —> co2 combines with RuBP to form two 3-PGA
captures co2 and adds carbon to RuBP by rubisco to make 3-pga then add energy to make G3P
chloroplasts
where photosynthesis occurs
plastids, located mostly in leaves
thylacoid
chlorophyll (photosynthesis pigment) is in thylacoid membrane
C3 plants
use the C3 photosynthetic pathway
a 3 carbon molecule (3-PGA) is first molecule produced
non-oxygenic photosynthesis
energy from sunlight, but electron donor is NOT water —> oxygen NOT released
cyanobacteria
first oxygenic photoautotrophs
adaptations for staying in euphotic zone
small size
spines, colonies to increase surface/volume ratio and drag
gas pockets
lipids deposits —> oils to help them float
adaptations for surviving adverse conditions (sinking out of photic zone)
resting stage in life cycle —> if they sink they make shell to remain alive in sediments
structural adaptions for desiccation: leaves
cuticle prevents evaporation: stomata let co2 inside leaf
h20 diffuses out of leaf in transportation
plants must balance water lost in transpiration with co2 absorbed for photo
adaptation for arid condition
stomata in pits —> trap moisture
ability to reduce leaf surface by curling
trade offs for reduced leaf area
capture less light
not shading neighbors
high energy cost of producing leaves with rigid cell walls
drought deciduous
plants lose leaves before annual dry season
winter deciduous
plants lose leaves before winter
trade off: need to grow leaves again next season
more roots than leaves
allocate more resources to roots (gather water) than leaves
mesic species
adapted to moist climates
xeric species
adapted to dry climates
photorespiration
catalyzed by rubisco
reaction of RuBP with o2 that releases co2
C4 plants
two kinds of photosynthetic cells
mesophyll cells where light reaction occur and co2 is fixed as 4 carbon acid and enzyme is PEP carboxylase
4 C is pumped into vascular cell and releases as CO2 for dark reactions
C4 as adapatation
compensates for high rates of photorespiration and carbon deficiency —> better for plants because don’t have to open stomata as much
photores rates increase w temp —> beneficial to have c4 pathway when temp increases
CAM pathway
crassulacean acid metabolism
same as c4 just not separated into two kinds of cell
net co2 intake
incorporation by photosynthesis - co2 release through respiration
light saturation point
photosynthesis increases with amount of PAR until it reaches a max
light compensation point
co2 uptake into photosynthesis balances co2 released in respiration
photoinhibition
photosynthesis often decreases at light levels beyond saturation
adaptations for low-nutrient environments
low absorption rate
high efficiency of nutrient use
lower growth rate —> slow growing
high ratio of leaves to stems —> deep roots
increased leaf longevity —> don’t shed leaves
animal cell vs plant cell
animal has much more proteins than plants → lots of nitrogen
plants have lots of carbs
grazers
herbivore
leafy material, especially grass
evolved from browsers
high crowned teeth
rumen
4 chambered stomach
browsers
herbivore
woody plants and their leaves
granivores
herbivore
seeds
only seasonally available in temperate and cold climates
frugiovers
herbivore
fruit
oxygen
needed to release energy contained in food
aquatic vertebrates —> oxygen
gills
countercurrent exchange maximizes oxygrn uptake from water
terrestrial animals in arid environments
fatty acids produce a lot of metabolic water —> fat in camels humps
estivation
summer formancy
diapause
stage of arrested growth in insects to survive dry periods
poikilothermic
animals have variable body temp influenced by envi
not much energy investment to change temp
ectothermic
homeothermic
animals maintain constant internal temp regardless of envi
lots of energy invested
endothermic
heterotherms
function either as ectotherms or endotherms depending on external circumstances
surface volume ration
decreases with size
influences most biochem and physiological processes as basic metabolism requires transfer of energy and materials between the organism's internal and external environments
daily torpor
dropping of body temp to ambient temp
endemic species
geographically restricted distribution
abundance
size of population
biomass
measure of abundance
population density
number of individuals per unit area
unitary organisms
zygote grows into genetically unique organism
module organisms
zygote grows into a unit of construction that then produces further similar modules (ramets or clones)
survivorship (lx)
number of individuals surviving to a given age as a proportion of original cohort size
age specific morality (dx)
difference between number of individuals alive for any age class and the next older age class
individuals died during given time interval
age specific morality rate (qx)
number of individuals dying during give time divided by number alive at beginning of that interval
individuals died in given age class
type 1 survivorship
convex
humans
type 2 survivorship
straight
adult birds, rodents, reptiles, many perennial plants
type 3 survivorship
concave
high mortality when young
fish, invertebrates, plants
fecundity table (fertility table)
mean number of females born in each age group
net reproductive rate (Ro)
number of females left during a lifetime by a newborn female
>1 pop is increasing
exponential growth
characteristic of populations inhabiting favorable envis at low population densities
demographic stochasticity
random variations in birth and death rates
environmental stochasticity
variation in envi can influence birth and death rates
allee effect
decline of a pop of small size
density dependent effects
influence pop in proportion to its size
density independent effects
influence a pop without regard to number of individuals
reproduction vs growth
allocation to reproduction reduces allocation to growth
asexual reproduction
offspring genetically identical to parents
benefits: all identical so adapted to envi and all able to reproduce
costs: no variation
binary fission
unicellular reproduce by dividing in half
parthenogenesis
give birth to all female offspring without fertilization
sexual reproduction
genetic variation but energetically costly
simultaneous hermaphrodites
both male and female —> male organ of one mated with female of another and vice versa and the same time
sequential hermaphrodites
individual will be male during one part of female during another
handicap principle
signals for mate are costly and may attract predators
indeterminate growth
continue to grow throughout adult hood
iteroparous reproduction
multiple reproductive events with fewer offspring
semelparous reproduction
single reproductive event with many offspring
optimal reproductive effort
balance between current and future reproduction that maximizes parent fitness
r strategy
many small offspring with little or no parental care
density independent growth limitation
k strategy
few larger offspring with considerable parental care
density dependent growth limitation
altricial
helpless at birth/hatching
precocial
more independent at birth/hatching
r plants
ruderal
live in high disturbance, low stress
c plants
competitive
live in low disturbance, low stress
s plants
stress-tolerant
live in low disturbance, high streess