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What is a biological community?
A level of biological organization made up of interacting populations of different species in the same area
Collective properties
Simple sum of all parts of a community
Examples of collective properties
Species richness; biomass
Emergent properties
Properties that are more than the sum of all parts
Examples of emergent properties
Niche partitioning; density-dependent predation; food web structure
Community structure
How a community looks
Community function
What happens in a community (processes)
Community composition
Which species are present
Tropical forest emergent layer
Tallest trees receiving the most sunlight
Tropical forest canopy
Main layer formed by tree crowns
Tropical forest undercanopy
Lower branches beneath the canopy
Tropical forest shrub layer
Small plants on the forest floor
Habitat structure factors
Carbon storage; light availability; age/size distribution; successional stage; microhabitats
Main drivers of habitat structure
Latitude; altitude; moisture
Guilds / functional groups
Groups of species that use resources in similar ways
Community functions
Nutrient cycling; carbon sequestration; erosion control
Local community assembly processes
Dispersal; abiotic factors; species interactions
Metacommunity
A group of communities that interact with each other
Clements view of communities
Communities act as tightly coevolved superorganisms
Gleason view of communities
Communities assemble loosely based on species tolerances and chance
Examples of bounded communities
Ponds; mangroves; bromeliads; tree-fall gaps
Why the tropics receive more energy
Earth's shape causes uneven heating concentrated at the equator
ITCZ
Low-pressure belt near the equator where trade winds converge and cause rainfall
Cause of wet and dry seasons in the tropics
Seasonal movement of the ITCZ
Why biodiversity is higher in the tropics
Less temperature variation and consistent solar energy
Effect of altitude on tropical temperature
Temperature decreases with increasing altitude
Orographic effect
Air rises over mountains, cools, condenses, and causes rain
Köppen climate maps
Classify biomes based on climate
Holdridge life zone system
Classifies communities using temperature, precipitation, and biotemperature
Biotemperature
Temperature as experienced by plants for photosynthesis
Potential evapotranspiration (PET)
Balance between water gained and water lost
PET greater than 1
More water lost than gained
Gross Primary Productivity (GPP)
Total energy captured by plants per unit area per time
Net Primary Productivity (NPP)
GPP minus respiration
Why NPP is hard to measure
Belowground growth, herbivory, and root exudates are difficult to quantify
Highly productive ecosystems
Coral reefs; algal beds; swamps; marshes; mangroves
Why tropical forests are "leaf factories"
Most productivity occurs in leaves rather than wood
Leaf litter in tropical forests
Dropped year-round in very large amounts
Dominant tropical soil types
Oxisols and Ultisols
Why tropical soils are nutrient-poor
Heavy weathering and leaching remove nutrients
Paradox of tropical luxuriance
High biomass and productivity on nutrient-poor soils
Where nutrients are stored in tropical forests
Plants and leaf litter rather than soil
Plant adaptations to low soil fertility
Shallow roots; more roots; tougher leaves; longer leaf lifespan
Importance of decomposition in the tropics
Rapid nutrient recycling maintains productivity
Key decomposers in tropical forests
Bacteria; fungi; termites; invertebrates; protists
Energy flow vs matter cycling
Energy flows through ecosystems; matter cycles within them
Why ecosystems are open systems
They exchange energy and matter with surroundings
Limitation on food chain length
Energy loss at each trophic level
Net Production Efficiency (NPE)
Efficiency of converting energy into biomass
Endotherms vs ectotherms
Endotherms lose more energy as heat
Bottom-up control
Plant defenses limit herbivore populations
Top-down control
Predators limit herbivores and protect plants
Trophic cascade
Chain reaction caused by removal of a species
Connectance
Actual interactions divided by possible interactions in a food web
Keystone species
Species with a disproportionately large effect on ecosystem structure
Typical tropical food chain length
Short (about four trophic levels)