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Biodiversity hotspots
areas with lots of species and diversity
island size
a geographic pattern of diversity, species richness increases logarithmicaly, w/ increase in land mass size
island
any kind of isolated habitat
latitude
a geographic pattern of diversity, species richness decreases linearly with an increase in latitude
ex. there are many lizards in areas near the equator, but only 2 in michigan, you lose species richness as you travel north
ecotone
a geographic pattern, a transition zone between 2 different types of habitat species richness, increases with convergence of different types of ecosystems
ex. estuary
equilibrium theory of island biogeography
proposed by Macarthur and Wilson, proposes that all islands reach an equilibrium of species richness, can be applied to other ecosystems. Can help explain S of islands of different sizes and distances from mainland.
S
equilibrium number of species, amount an island can support balanced by extinction and immigration
is the carrying capacity for the community
(draw an isocline graph for S)
an isocline graph
graphs 2 things at once
if there are no species on an island, the extinction rate is
0
land bridge
a type of island, forms when sea level rises or from other changes, isolating the island from the mainland
begins with species richness of mainland, loses the richness over time since the island is smaller (came from a bigger habitat and got divided up)
(draw a graph)
relaxation fauna
species that remain isolated on the new island, they are the ones that don’t die out and remain when S is reached
oceanic
type of island, there wasn’t one before and it appears, typically forms from volcanic activity
begins with 0 richness, gains it over time
ex. Hawian islands
(draw graph)
islands closer or farther from mainland will have a higher S
closer, there is a higher immigration rate due to shorter traveling distance
(draw graph)
farther islands will catch up eventually in immigration, however never in population size because they have the same rate of extinction as the closer islands. Therefore farther islands always have a lower S
however farther islands usually have more unique species
bigger or smaller island will have a higher S?
bigger, species are more likely to encounter a big island than a small island, and a bigger island can support more species than a smaller one
(draw a graph)
Significance of mangroves
mangrove trees are kinds of islands, provide a land habitat in the middle of a body or water that many insects live on. Scientists killed all the species on a mangrove and essentially made it into an oceanic island. Species richness over time then came back up, similar to what it was before scientists killed the species.
shows almost any isolated habitat follows island biogeography theory
fragmented habitats
a habitat that was once large and continuous but now is broken into smaller, more isolated patches. Roads and national parks create them, as well as natural disasters and environmental obstacles. They are concerning for endangered animals especially, as the habitat decreases in S as it becomes fragmented and access to resources decreases
one way to bring fragmented habitats together
to make land bridges over roads, expands habitat and resource access, as well as helps preserve gene flow and diversity
ecological niche
specific role of a species in an ecosystem
includes all physical, chemical, and biological components needed for that species’ survival
physical resources
abiotic components that must be assimilated for survival, including O2, CO2, water, light (for photosynthesizers), nutrients, and minerals
physical factors
abiotic parameters that affect where an organism can live including temp, pH, salinity, and light
draw a graph of a single species with no competition for prey size vs. individuals eaten
looks like a bell shaped curve
niche width
quantifies the variance in resource use. The more adaptable the species, the broader the niche. Specialists have less width, generalists have more
specialist
can live in few habitats, eat few types of food, tolerate a narrow range of environmental conditions, K selected
generalists
can live in many habitats, eat many types of food, tolerate a wide range of environmental conditions, R selected
Hutchinsonian niche
quantifies the level of resources and factors that a species needs for survival, you can plot several resources and factors in multi-dimensional space.
Can plot multiple single factor niches so they become multi factor niches
can measure for example prey size eaten, oxygen content of water lived in, water salinity lived in. Turns the graph into a 3d box
niche multidimensional hypervolume
plots of multiple resources and factors and the range within which a species is found
has unlimited dimensions
shows Hutchinsonian niche, quantifying ecological niches
used in ecology to see if conditions are appropriate to be able to reintroduce endangered species to a habitat, or see why they are endangered in the first place
niche separation
distance between mean resource curves for 2 species. There is sometimes overlap and sometimes not, the curves may intercept or not

niche overlap
area of resource curves that overlap, the overlap is where competition occurs

competition
occurs when 2+ niches overlap in 2+ species, leads to a decrease in population of at least 1 species’ population because there are less resources available , is a kind of interaction
even if there is plenty of food, they can fight over it, creating competition
preemptive competition
competition for specific space where resources can be obtained
not the same as territoriality
ex. plants want to be in the best sunlight spot, compete but not necessarily intentional
interference competition
once species limits anothers access to a resource even if its abundant
can have to do w/ territoriality
ex. bears picking the best spot on the river to catch salmon
lions and hyenas hating eachother and not wanting to share territory
butterflies fight over abundant nectar sources
some trees poison the ground so others cannot grow there
allelopathy
producing chemicals to interfere with another plant’s ability to grow
exploitation competition
once species uses more of a limited resource, leading to the detriment of the other species
1st species is usually better at exploiting
3 competition types
interference, preemptive, exploitation
competition isocline
predicts the result of competition between 2 species, works best with exploitation competition where there is a clear competition and resource, but there are multiple kinds: competitive exclusion, weak competition/stable equilibrium, and unstable equilibrium
(draw graphs of each)
6 elements of a competition isocline
N1 pop size of species 1
N2 pop size of species 2
a12 effect on species 1 by 2
a21 effect on species 2 by 1
K1 carrying capacity of species 1
K2 carrying capacity of species 2
competitive exclusion isocline
if there is a larger difference in population between the two species, there will be less competition. Always eventually ends with the species with the larger K outcompeting the other. If the dominant species is at its K, the other species population is at zero. Therefore K is the amount it takes to bring species 2 to zero
species 1 is better at exploiting resources
draw graph
competitive exclusion
local extinction or near extinction of a species due to interspecific competition
species with a higher K, better exploitation, etc. will outperform others
the species with the highest isocline is the “winner”
weak competition/stable equilibrium isocline
no species will die out, competition is less intense. Will always over time acheive an equilibrium in which there are steady populations for both species, and their Ks are similar. Alpha values are smaller than other forms.
can lead to character displacement over evolutionary time
competition still has an effect but not exclusion.
Character displacement
natural selection essentially, separation of 2 species in morphology or behavior as a result of competition. Occurs over evolutionary time, an overlap will evtually separate. Occurs all the time in the tropics and other table environments
ex. wildabeest, zebra, and giraffess have the ability to eat all the same plants, but eat different ones at different heights, showing stable equilibrium
ex. parasites
ex. warblers eating at different tree heights
draw graph
unstable equilibrium isocline
K’s are roughly equal, but are above alpha values. It is possible for one species to be driven no population, but also possible for equilibirum to be acheived depending on the initial population sizes. Competition is fairly intense.
unstable equilibrium
occurs when the result of competition depends on initial population sizes. Alpha values are large, bigger than K. Can lead to either competitive exlcusion or character displacement
unequal population sizes leads to competitive exclusion
explain how the population of asian tiger and native mosquitoes relate to isoclines
the asian mosquitoes were brought to the US in a tire shipment, which held a lot of water. Native mosquitoes can control the asian populations through competition. In wilderness and trees, the two populations show an unstable equilibrium isocline. In tires, however, competitive exclusion occurs and the asian mosquitoes outcompete.
producer
made food with photosynthesis, are autotrophs
ex. plants, algae, some bacteria
consumer
also known as a predator or heterotroph, obtain food from eating living organisms
everything but producers
primary consumers
eat producers, are herbivores
secondary consumers
eat primary consumers, are carnivores
ex. praying mantis, whale
tertiary consumer
eat secondary consumers
omnivore
eat producers and consumers
parasite
an organism that feeds on another while it is living, or inhabits the host organism
the host is rarely killed
also known as micropredators
parasitoid
larvae of an organism parasitizes their host, but never the adults
frequently kills the host
only in flies and wasps
detrivore
eat waste products from other organisms like dung and cast off body parts
heterotrophic
ex. dung beetle (NOT vultures)
decomposers
not detrivores,breaks down organic matter into simpler compounds
recycles nutrients and minerals back into soil and water
ex. some fungi and bacteria
prey defense mechanisms
urtication, camoflauge, escape, and chemical sequestration
urtication
irritating predators by physical structures on the body that poke you
ex. porcupine
chemical sequestration
when an organism stores toxic chemicals
tobacco, milkweed (butterflies consume it and can become toxic)
Aposematic coloring
coloration to warn that something is toxic, usually red or yellow
ex. poison arrow frogs: consume toxic insects, become toxic to predators
poison dart frogs, toxic to the touch. Red and yellow
optimal foraging theory
organisms will obtain the max amount of energy under a given set of foraging conditions
finding a balance between calories and time
want the most calories
how much can you eat vs. how hard you have to work
includes energy maximizers and time maximizers
ex. wolves could eat mice, but a moose has many more calories, but also requires more energy to catch
energy maximizer
species that make receiving the most energy per meal their priority, want the most food they can eat
snakes, angler fish
time maximizer
species that make spending the least amount of time catching a meal their priority
grazing species like rabits or cows
different respinses to prey increase
functional response, numerical response, combined responsefu
functional response
each individual predator consumes more prey individuals, leads to healthier individuals of the predator population, when there is more food to eat, individuals will eat more
numerical response
predators increase in numbers, overall predator population consumes more prey, not individuals
breeding more, not eating more, causes an overall increase in population
strongest functional response has the lowest numerical response
combined response (to prey increase)
if predators eat or breed more, the % of predation increases
predation is density dependant up to a certain point, because you can only eat so much
prey can always outbreed predators
ex. all wildabeest give birth at the same time, lions have tons of food, but they can only eat so much (can’t eat everything)
competition in a trophic guild can lead to stable equilibrium which can then lead to charatcer displacement
ecological guild
group of ecologically similar organisms, they compete if they are in the same environment. Can be in a guild even if they are not in the same envrionment
allopatric
indiviauls of 2 species not living within the same community
no competition
ex. deer and moose (moose go crazy from deer brainworm)
sympatric
2 species living in the same community
ex. Lake Malawi: 250 sympatric species of cichlids, has 10 distinct trophic guilds, leading to many super specific niches and a complex environment
community equilibrium theory
every species and combination of species has a limit to their population size (equilibrium) within an ecosystem
competition and predation are main community structuring forces
interspecific competition
component of community equilibrium theory, species within trophic guilds competing for scarce resources structure communities
draw back to fundamental nice: Hutchinsonian niche for species in absence of competition
but with competition: realized niche and character displacement→subset of fundamental niche actually utilized by species
competition decreases realized niche
predation
other component of equilibrium theory, predators consume prey and its corresponding effects structure communities
if you remove starfish from shore, muscles wipe out all species in the area
removing a predator can do more than impact population of prey
keystone species
a predator that affects an ecoystem to a far greater degree than their biomass would suggest
removal can effect many seemingly unrelated species
ex. kangaroos, rats, starfish, beaver, wolves
NOT termintes, have a lot of biomass
trophic cascade
removal of a keystone species causing changes to multiple trophic levels

ex. when elk populations increase, they eat more vegetation, causing loss of trees for songbirds and beavers
community nonequilibrium theory
species and combinations of species are rarely at equilibrium, de-emphesizes K
communities are far more dependent on abiotic, density indep. effects for their structure
abiotic extrinsic view
richness, diversity, and complexity increases in favorable environmental conditions
will suffer under disturbances like floods, droughts, fire, a