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Biodiversity
The variety of life in an ecosystem measured at the genetic, species, and ecosystem levels
Genetic diversity
The variety of genes and genetic differences among individuals within a population of the same species
Species diversity
The variety of species in an ecosystem and the balance of their population sizes
Ecosystem diversity
The variety of habitats and ecosystems within a given area
Species richness
The total number of different species in an ecosystem
Species evenness
How evenly individuals are distributed among the different species in an ecosystem
High biodiversity
A condition that generally increases ecosystem health, resilience, and the ability to withstand disturbances
Genome
The complete set of genetic information in an organism
Mutation
A random change in DNA that can create new genetic variation
Genetic recombination
The rearrangement of genetic material that creates new combinations of genes in offspring
Genetic variation and adaptation
Greater genetic variation increases the likelihood that some individuals possess traits that help them survive environmental changes
Population bottleneck
An event that drastically reduces population size and usually decreases genetic diversity
Bottleneck effect
The loss of genetic variation when a small surviving population no longer represents the original population's genetic diversity
Inbreeding
Reproduction between closely related individuals
Inbreeding depression
Reduced fitness caused by increased expression of harmful genetic mutations in offspring of related individuals
Ecosystem resilience
The ability of an ecosystem to recover its original conditions after a disturbance
Ecosystem services
Natural goods and processes that benefit humans economically or by supporting life
Provisioning services
Ecosystem goods obtained directly from nature, such as food, timber, and medicinal resources
Regulating services
Ecosystem processes that moderate environmental conditions, such as climate regulation and air purification
Supporting services
Natural processes that support human activities and make them easier or less costly, such as pollination
Cultural services
Ecosystem benefits associated with recreation, tourism, and scientific knowledge
Carbon sequestration
The capture and storage of carbon dioxide, including by trees through photosynthesis
Pollination
The transfer of pollen that enables plants to reproduce and can increase agricultural production
Economic consequences of biodiversity loss
Reduced ecosystem services can cause crop failures, lost fishing income, higher healthcare costs, and increased environmental damage
Island biogeography theory
A theory explaining how island size and distance from a mainland influence the number of species present
Island size and species richness
Larger islands generally support more species because they provide more habitats, resources, and ecological niches
Island distance and species richness
Islands closer to the mainland generally support more species because colonization is easier
Colonization
The establishment of a species in a new habitat
Habitat island
An isolated patch of natural habitat surrounded by a different environment, often created by human development
Habitat fragmentation
The division of a large, continuous habitat into smaller, disconnected patches
Wildlife corridor
A strip of habitat that connects isolated habitat patches and allows organisms to move between them
Benefits of wildlife corridors
They support migration, gene flow, recolonization, and movement between habitat patches
Edge effect
Abrupt changes in biotic and abiotic conditions near the boundary between two different habitats
Habitat patch size
Larger habitat patches generally contain more interior habitat and can better support species that require large territories
Habitat fragmentation and biodiversity
Fragmentation often reduces biodiversity by isolating populations, reducing habitat area, and increasing edge effects
Adaptive radiation
The evolution of one ancestral species into multiple species adapted to different ecological niches
Ecological niche
The role of an organism in its ecosystem, including its resource use and environmental requirements
Ecological range of tolerance
The range of environmental conditions an organism can endure before injury or death occurs
Optimal range
Environmental conditions under which an organism can survive, grow, and reproduce successfully
Zone of physiological stress
Environmental conditions under which an organism survives but experiences reduced growth, activity, or reproductive success
Zone of intolerance
Environmental conditions outside an organism's survival limits that result in death
Indicator species
Species that are sensitive to environmental changes and provide evidence about ecosystem health
Characteristics of indicator species
They often have narrow tolerance ranges and respond quickly to pollution or environmental disturbance
Examples of aquatic indicator species
Stonefly nymphs, mayfly nymphs, caddisfly larvae, and trout
Water temperature
A water-quality parameter that affects aquatic organisms and the amount of dissolved oxygen water can hold
Dissolved oxygen
Oxygen present in water that aquatic organisms require for cellular respiration
Low dissolved oxygen
A condition that can cause physiological stress, suffocation, and death in aquatic organisms
Water pH
A measure of acidity or alkalinity that affects aquatic organisms and the toxicity of dissolved metals
Low pH and metal toxicity
More acidic water can make dissolved metals more toxic to aquatic organisms
Eutrophication
Nutrient enrichment of water that can cause algal blooms and oxygen depletion
Algal bloom
Rapid growth of algae that can contribute to low dissolved oxygen when the algae die and decompose
Natural disturbance
A natural event that disrupts the structure or function of an ecosystem
Examples of natural disturbances
Tornadoes, hurricanes, droughts, forest fires, earthquakes, volcanic eruptions, and asteroid impacts
Periodic disturbance
A disturbance that occurs at regular intervals, such as seasonal changes between wet and dry periods
Episodic disturbance
A disturbance that occurs occasionally at irregular intervals, such as a hurricane or drought
Random disturbance
A disturbance without a regular pattern, such as an earthquake or volcanic eruption
Natural climate change
Long-term climate variation caused by natural processes, including changes in Earth's orbit and tilt
Natural disturbances and biodiversity
Major disturbances can reduce biodiversity by destroying habitat and pushing organisms outside their tolerance ranges
Migration
The movement of organisms to a different habitat, sometimes in response to environmental changes
Primary succession
Ecological succession that begins on bare rock or another surface without established soil
Secondary succession
Ecological succession that begins where established soil remains after a disturbance
Primary succession examples
Colonization of newly formed volcanic rock and rock exposed by retreating glaciers
Secondary succession examples
Recovery after a forest fire, flood, or deforestation when soil remains
Pioneer species
Early successional organisms that establish themselves first and tolerate harsh environmental conditions
Primary succession pioneer species
Mosses and lichens that can grow on bare rock and help initiate soil formation
Secondary succession pioneer species
Fast-growing grasses, wildflowers, and other plants that establish in existing soil
Middle successional species
Plants such as shrubs and fast-growing trees that establish after pioneer species improve soil conditions
Late successional species
Long-lived, often shade-tolerant trees that establish after soil becomes deeper and more nutrient-rich
Climax community
A relatively mature community associated with later stages of succession
Why primary succession takes longer
Soil must develop before many plants can establish, whereas secondary succession begins with existing soil
Soil formation during primary succession
Mosses and lichens weather rock, while decaying organic matter contributes to initial soil formation
Natural selection
The process by which individuals with advantageous heritable traits tend to survive and reproduce more successfully
Adaptation
A heritable trait that increases an organism's ability to survive and reproduce in a particular environment
Fitness
An organism's ability to survive and reproduce
Selective pressure
An environmental condition that affects survival and reproductive success
Evolution
Changes in the heritable characteristics of populations across generations
Environmental change and adaptation
Environmental changes can make previously beneficial traits disadvantageous and favor different traits
Finch beak adaptation
Finches with beaks better suited to available food can survive and reproduce more successfully
Genetic diversity and environmental change
Greater genetic diversity increases the probability that a population contains individuals able to survive new environmental conditions
Rate of environmental change
Rapid environmental changes can cause population declines when species cannot adapt or migrate quickly enough
Specialist species
Species with narrow habitat, food, or environmental requirements that are often more vulnerable to disturbance
Generalist species
Species able to use a wider variety of resources and habitats, often making them more adaptable to environmental change
Endangered species
Species facing a high risk of extinction
Overexploitation
Harvesting or killing organisms faster than their populations can replenish themselves
Poaching
Illegal hunting, capture, or killing of wildlife
Invasive species
Non-native species that establish and spread in a new area and cause ecological or other harm
Exotic species
A species introduced outside its native range; not every exotic species becomes invasive
Characteristics of invasive species
They may reproduce rapidly, compete aggressively, adapt easily, and lack effective natural predators
r-selected species
Species that generally reproduce rapidly, produce many offspring, and provide relatively little parental care
Biotic potential
The maximum reproductive capacity of a population under ideal conditions
Invasive species and native species
Invasive species can reduce native populations by competing for food, water, sunlight, and space
Why generalists often become invasive
Their flexible diets and broad habitat requirements help them survive in new environments
Why specialists are often endangered
Their specific habitat and food requirements make them less adaptable to environmental change
Interspecific competition
Competition for resources between members of different species
HIPPCO
Habitat loss, invasive species, population growth, pollution, climate change, and overexploitation as major threats to biodiversity
Habitat loss
The destruction or conversion of natural habitat for activities such as agriculture, roads, and urban development
Pollution and biodiversity
Pollutants such as oil and pesticides can injure or kill organisms and reduce populations
Climate change and biodiversity
Changes in temperature and precipitation can shift habitats faster than some species can adapt or migrate
Population growth and biodiversity
Increasing human populations can increase habitat destruction through urbanization and agricultural expansion
CITES
An international agreement through which countries regulate international trade in endangered species