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spherical body symmetry
any plane through body divides it in mirrored halves
good for floating and rolling
rare in animals, common for single celled organisms
radial body symmetry
2 planes through centre divids body into mirrored halves
biradial
only 2 planes produce mirroed halves
bilateral body symmetry
right and left mirroe halves along sagittal plane
great for forward movement
allows for head region
germ layer
distinct layer of cells that develop shortly after fusion of gametes
all tissues and organs develop from these
dipleblasts
2 germ layers
ectoderm and endoderm
triploblasts
3 germ layers
ectoderm, endoderm, mesoderm (muscles and curculatory system)
blastula
early cell divisions form a hollow ball of cells
deuterostomes
blastopore becomes anus
radial cleavage
protostomes
blastopore becomes mouth
spiral cleavage
body cavity (coelom)
internal space, tissues layers and embryonic development
allows organs to not be attached to outer wall (allows for movement)
schizocoely
division in mesoderm becomes coelom
schizocoely
enterocoely
cavity forms right away
deuterostomes
ecdysozoa
clade in the protostomes
includes those with a cuticle that is shed then replaced
lack cilia
embryos do not develop by spiral cleavage
spiralia
clade in the protostomes
animals developing mostly from spiral cleavage
panarthropoda
unranked clade in eudyzoa including phylum arthropoda, tardigrada, and onychophora
animals that molt, have segmented bodies, paired appendages with claws, and a brain
phylum arthropoda
segmented body (segments often fused to form tagmata)
hard outer cuticle made from protein and a polysaccaride called chitin
jointed appendages
arthropoda circulatory system
open circulatory system
fluid (haemolymph) pumped by heart into open spaces that surround organs and tissues
arthropoda gas exchange
across the body integument (outer covering) in some
others have gills or tracheal systems
subphylum crustacea
all appendages exept first antennae are biramous
2 pairs of anntenae
nauplius larvae
crustacea respiratory system
large crusts use featherlike gills
small crusts lack gills and have passive transfter across thin sections of body wall
terrestrial isopods have thin structure on ventral posterier that allows gas transfter (needs moistue)
crustacean circulatory systen
open system
hemolymph that contributes to gas transfer
crustacean excretory system
anterior to esophagus
called green glands
hydrostatic pressure of hemocoel forces fluid to filter into gland
resorption of salts and amino acids occurs as filtrate passes the excretory tubule and bladder
crustacean nervous system
fused ganglia throughout body
brain connects to eyes and 2 pairs of antennae
subesophageal ganglion connects to brain and suplies nerves to body
crustacean sense organs
compound eyes and statocycts (balance and equilibrium)
tactile bristles (setae)
remipedia
crustacean
primitive vermaform
predator
cephalocardia
crustacean
benthic marine detritavore
lack carapace, has cephalic sheild (head)
no eyes
branchiopoda
crustacean
mostly freshwater
may have carapace, appendages phyllopodous (leaf like)
many produce drought resistant eggs of zygotes
malacostraca
crustacean
may have carapace
body 19 or 20 segments
6-8-6 somite makeup
subclass eumalacostraca (order decapoda)
well developed carapace enclosing gill chamber
3 pairs of maxillipeds, 5 pairs of pereopods
one or more anterior pereopods are chelate (clawed)
carcinization
evolution towards carb like morphology
shortened abdomen
theostraca
branacles
reduced adult morphology
free-living and parasitic examples
hexapoda
6 legs
all legs uniramous (segments attach end to end without splitting)
3 body segments (tagmata) head thorax and abdomen
order collembola
class entognatha
small and terrestrial
no spiracles or thracheae (gas transfer across body wall)
insect larvae
true legs only on thorax
fleshy prolegs can occur on abdomen
metamorphasis
characteristic of clade ecdysozoa (common in arthorpods and hexapods)
exoskeleton shed between stages
ecdysis
molting
instar
each life stage between molts
ametabolous metamorphasis
slight changes or increase in size
found only in primitive wingless orders
hemimetabolous (incomplete) metamorphosis
gradual change
young are called nymphs (wingless)
wing pads in early instars where adult wings will grow
egg → nymph→adult
grasshopper, dragonfly
holomatabolous (complete) metamorphosis
about 88% of insects
larvae and adults often live in different environements and have different morphology
egg→larvae→pupa→adult
arthropoda subphyla
crustacea (lobsters, crabs)
hexapoda (insects)
myriapoda (centipedes, millipedes)
chelicerata (spiders, horseshoe crabs)
trilobita (extinct!!)
myriapoda
2 part body structure (head and multisegmented trunk)
uniramous multiarticulate appendages
head includes antennae, mandibles and maxillae
chelicerata
spiders, scorpions, ticks
2 part body plan: prosoma (7 segments) and opisthosoma (up to 12 segments+telson)
specialized appendages (chelicerae, pedipalps and 4 pairs of walking legs, antannae are absent)
class euchelicerata- subclass merstomata
modern horsehoe crab nearly unchanged from ancestors
unsegmented carapce covers body infront of broad abdomen and telson
5 pairs of walking legs
gills on ventral abdomen
carapace have 2 compound and 2 simple eyes
subclass arachnida
scorpians, spiders, ticks, mites
2 tagmata (prosoma and opisthosoma)
prosoma has carapace, pair of chelicera, pair of pedipalps, and 4 pairs of walking legs
opisthosoma lacks appendages
no antenna or mandibles
arachnida-acari (subclass arachnida)
ticks and mites
not monophyletic
complete fusion of body segments
anterior portion is caputulum, remaninder is idiosoma
family ixodide (ticks) (subclass arachnida)
hard ticks
sclerotized dorsal covering (scutum)
feeds on hosts for days to weeks, may carry disease
blood feeding ectoparasite with chelicerae for cutting skin
egg→6-legged larvae →8-legged nymph →adult
black legged tick (lyme disease) and dog tick in ontario
family argasidae (ticks) (subclass arachnida)
soft ticks, lack scutum
feed for short periods (>1 hour) may carry disease
ablypygi (subclass arachnida)
whip spiders
carapace, narrow pedicel, no telson, chelicerae specialized as fangs, pedipals raptorial
1st pair of appendages extented and sensory
uropygi (subclass arachnida)
whip scorpions and vinegaroons
carapace, whip-like telson (sensory), first walking legs elongated (sensory)
repugnatorial glands near anus spray acid at attackers
aranae (subclass arachnida)
carapace, chelicarae specialized as fangs (some have vemon), pedipaps modified as male copulatory organs
spinnerets produce silk
spider webs
fiborous protein
up to 4 pairs of abdominal spinnerets
producing web is energetically costly
some spiders use for catching prey others dont
scorpiones ((subclass arachnida)
carapace like sheild
3 body section (prosoma, mesosoma, metasoma)
large chelate pedipalps
spine-like telson with poison gland
pseudoscorpiones (subclass arachnida)
small
segmented carapace-like shield
chelicerae chelate with silk glands
phylum onychophora
ecdysozoa, evolutionarily between nematodes and arthropods
similar to arthropods, body cavity is a hemocoel divided into sinuses
cannot close spiracles, tracheal system evolved independently from arthropods
predaceous velvet worms
phylum nematoda
unsegmented roundworms
cuticle secreted by hypodermis, rigid and thick, made of collagen, must be molted
no circular muscles (only longitudinal) whip like movement
parasites, some free living (Caenorhabditis elegans)
complete gut, waste expelled by pseudocoelom pressue
gas exchange across body wall
nematode reproduction
amoeboid sperm crawls to female (sperm lack flagella)
Ascaris lumbricoides
intestinal roundworm
largest nematode infecting humans
female worm lays 200000 eggs a day
adults live in intestine, eggs released in feces
juveniles migrate through body
Dirofilaria immitis
dog heartworm
transmitted by mosquito bite
adults live in vessels around heart
annelida
segmented roundworms
secreted nonchitinous outer layer
annelid organ systems
complete digestive system
closed circulatory systems
complex nervous system
Eucoelomates
meristem(annelids)
independent movement of each segment
Subphylum Aclitellata: Class Polychaeta
paddle worms
mostly marine
paired paddle-like appendages called parapodia with chitinois hairs for movement
subphylum aclitellata: class polychaeta
sedentary forms live in tubes, suspension or deposit feeders
reporudce sexually by relasing eggs and sperm into water (oviparous)
marine forms have trochophore larvae, freshwater NO larval stage
subphylum chitellata: class oligochaeta
4 bundles of chaetae on each segment (few)
no larval stage
subphylum chitellata: class hirudinea
leeches
34 segments, anterior sucker and posterior sucker
ocelli (eye spots)
predatory leeches are nocturnal and negatively phototactic
subphylum chitellata: class branchiobdellida
ectosymbionts of crayfish
similar to leeches but only 11 segments and no anterior sucker
errantia
mostly free-moving annelids
traditionally called polychaetes
well differentiated head
parapodia sense organs