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Option C: Ecology and Conservation

C1 Species and Communities

  • Survival factors

    • U: The distribution of species is affected by limiting factors

      • Limiting factor = a component of an ecosystem which limits the distribution or numbers of a population

        • Defines optimal survival conditions according to its effect on a species when in deficiency or excess

        • Can be biotic or abiotic

          • Biotic factors include interactions between organisms

            • Either intraspecific (within a species)

            • Or interspecific (between species)

          • Abiotic factors include environmental conditions

            • E.g. light, temperature, salinity (concentration of salts in water or soils), rainfall, wind velocity, soil pH, etc.

        • Typical abiotic limiting factors for plants within a community

          • Temperature

            • Plants can only survive within a narrow range of temperatures to which they are adapted

            • High temperatures will increase the rate of water loss by evaporation and may also denature metabolic enzymes

            • Low temps may cause plant sap to freeze

              • The expansion of frozen water in xylem can cause trunks to split

            • Certain species of woody plants synthesis antifreeze protein to prevent crystal formation in frozen cells

              • E.g. maple trees

          • Water availability

            • Water is needed for photosynthetic processes + maintaining cell turgor

            • Xerophytes are plant species that are adapted to survive in dry and arid environments such as deserts

              • E.g. cacti

            • Hydrophytes are plant species that are adapted to survive in frequently waterlogged soils

              • E.g. rice

          • Light availability

            • Light is essential to the process of photosynthesis whereby plants produce organic molecules

            • Low-growing plants will typically possess darker leaves in order to optimize their light conversion

              • Dark = more chlorophyll

            • Certain seaweeds have pigments adapted to absorbing blue wavelengths

              • E.g. kelp

              • Red does not easily penetrate water

        • Typical abiotic limiting factors for animal within a community

          • Temperature

            • Animal survival typically relies on narrow temperature range

              • Few animals can survive temperature extremes

            • Poikilotherms cannot maintain thermal homeostasis and must occupy environments according to temperature needs

            • Homeotherms can regulate their own internal body temperatures and hence can occupy a wider range of habitats

            • Body size will play a fundamental role in determining an animal’s capacity to retain heat

              • The surface area to volume ratio

          • Territory

            • Territorial boundaries may determine an animal's capacity to attract mates, rear young, forage for food and avoid predators

            • Territories may be temporary (e.g. migration, breeding sites) or alternatively may be permanent

            • In certain species, juveniles may have different environmental requirements to adults

              • E.g. tadpoles and frogs

            • Establishment of territories can lead to intraspecific and interspecific competition

          • Food availability

            • Availability of food supply is a critical determinant in influencing population size and distribution

            • Animals may require a particular plant or animal species as a food source

              • This limits their spread to certain regions

              • Seasonal or geographic variations may directly affect food availability within a population

                • E.g. seasonal migrations

      • Law of Tolerance

        • The Law of Tolerance was proposed by Victor Ernest Shelford (American zoologist) in 1911

        • States that populations have optimal survival conditions within critical minimal and maximal thresholds

          • When populations are exposed to the extremes of a particular limiting factor (too much or too little), the survival rates drop

        • The distribution of species in response to a limiting factor is a bell-shaped curve with 3 distinct regions

          • The optimal zone

            • Central portion of a curve which has conditions that favor maximal reproductive success and survivability

          • Zones of stress

            • Regions flanking the optimal zone where organisms can survive but with reduced reproductive success

          • Zones of intolerance

            • Outermost regions in which organisms cannot survive

            • Represents the extremes of the limiting factor

          • Example:

    • A: Distribution of one animal and one plant species to illustrate limits to tolerance and zones of stress

      • Plant example

        • Plant growth varies greatly in response to salinity levels (concentrations of salt within the soil)

        • Plants that are not salt tolerant are called glycophytes

          • They are easily damaged by high salinity

          • Most plant species are considered glycophytes

        • Plants that are salt tolerant are called halophytes

          • They may become stressed in freshwater environments

          • Less than 2% of all plants are halophytes

        • Cultivation of land for agriculture causes the water table to rise and concentrates salt at the roots

          • Such cultivation includes irrigation and grazing

          • This makes it harder for glycophytes to extract water from the soil and the uptake of salt can be toxic to the plant

        • Understanding salt tolerance for different plant species is therefore critically important to effective crop farming

      • Animal example

        • Coral species form connected reefs → these are greatly impacted by changes in oceanic temperature

        • Coral polyps receive nutrition from photosynthetic zooxanthellae that lives within the polyp’s endodermis

          • Zooxanthellae = an algae

          • Endodermis = an inner layer of cells in the cortex of the root

        • The Zooxanthellae cannot survive in lower ocean temperatures

          • Below 18 degrees Celsius

        • Increases in ocean temperature cause Zooxanthellae to leave the coral tissue → this leads to coral bleaching

          • Above 35 degrees Celsius

        • Reef-building coral species therefore have a typical optimal growth range in temperate waters between 20 and 30 degrees Celsius

          • This correlates to the tropical and sub-tropical regions of the world (near the equator)

  • Species distribution

    • S: Use of a transect to correlate the distribution of plant or animal species with an abiotic variable

      • Quadrats and transects can measure the distribution of a plant or animal species in response to an incremental abiotic factor

        • Incremental = happening gradually

        • Quadrats = rectangular frames of known dimensions than can be used to establish population densities

        • Transects = a straight line along an abiotic gradient from which population data can be recorded to determine a pattern

        • Quadrats and transects are used together

      • Quadrats are placed at regular intervals along the transect line to generate population data

        • Basically, placing a quadrat down regularly and observing/counting the difference in distribution of X species based on the change of Y abiotic variable

        • The quadrats will show the changing distribution pattern of a species in response to a change in an abiotic variable

        • This data can be used to identify optimal conditions + zones of stress and zones of tolerance

      • Sampling with Quadrats and Transects

        • Quadrat counting method: count all variables mostly inside the quadrat (not mostly out)

        • Line transects: find a line, place quadrat every x meters

        • Transect data

          • Transects are used to assess species distribution in correlation with any abiotic factor that varies across a measurable distance

            • These factors could include elevation, elemental exposure, temperature, light levels, pH, humidity and more

        • A kite graph with the transect data can be used to represent changes in species distribution in a clear and effective fashion

          • The relative width of each kite represents the abundance of an organism at a particular point along a transect

          • Example:

  • Ecological niche

    • U: Each species plays a unique role within a community because of the unique combination of its spatial habitat and interactions with other species

      • Ecological niche = the functional position and role of an organism within its environment

        • Consists of all physical and biological conditions which determine the organism’s survival and reproductive prospects

        • Is comprised of various components, including:

          • The habitat in which an organism lives

          • The activity patterns of the organism

            • E.g. periods of time during which it is active

          • The resources it obtains from the environment

            • E.g. food sources, territorial boundaries, etc.

          • The interactions that occur with other species in the community

            • E.g. predator prey relationships, competition, etc.

    • U: Two species cannot survive indefinitely in the same habitat if their niches are identical

      • If two distinct species share an identical niche, there will be interspecific competition for available space and resources

        • This competition will result in the fitness of one being lowered by the presence of the other

        • The less well-adapted species will struggle to survive and reproduce

          • Eventually it will be eliminated from the niche

      • Interspecific competition within a shared niche will typically prompt one of two responses

        • Competitive exclusion → one species uses the resources more efficiently, driving the other species to local extinction

        • Resource partitioning → both species alter their use of the habitat to divide resources between them

          • Niche separation

      • Niche differentiation diagram:

    • S: Analysis of a data set that illustrates the distinction between fundamental and realized niche

      • Some species may not be able to occupy their entire niche due to the presence or absence of other species

        • Hence, a species may occupy a small subset of their niche than is theoretically possible

        • This creates fundamental vs. realized niche

      • Fundamental niche = the entire set of conditions under which an organism can survive and reproduce (where it could live)

        • The theoretical habitat

        • Example: in a rocky shore environment, the Chtalamus barnacle can potentially occupy the entire rocky shore, if in isolation

      • Realized niche = the set of conditions used by an organism after including interactions with other species (where it does live)

        • The actual habitat that is completely occupied by an organism in the presence of competing species

        • Example: the Chtalamus barnacle only occupies regions where the Semibalanus barnacle is absent

      • Diagrams:

        • Niches

        • Experimental data

        • Types of barnacles

  • Species interactions

    • U: Interactions between species in a community can be classified according to their effect

      • In nature, no species exist in total isolation → all organisms interact with both the abiotic environment and other organisms

        • The interactions between species can be classified according to their effect on the organisms involved

      • Herbivory

        • The act of eating only plant matter

          • That’s why primary consumers are called herbivores

          • Herbivores ay employ different feeding strategies

            • E.g. mucivores feed on plant sap while granivores feed on seeds, etc.

        • Herbivory can be either harmful or beneficial to plant species as a whole

          • Harmful: Certain types of beetle may feed voraciously on the leaves which causes crop failure

          • Beneficial: Fruit-eating animals spread the seeds from a fruit in their feces, which promotes overall seed dispersal

        • Examples of herbivores: Rabbit, cow

      • Predation

        • A biological interaction whereby one organism hunts and feeds on another organism

          • Predator and prey

        • The dependance of the predator on the prey as a food source intertwines their population levels

          • If the prey population drops, the predator numbers will fall as well as intraspecific competition has to increase

          • If the prey population rises, predator numbers will increase as a result of the ober-abundance of a food source

            • No intraspecific competition necessary = even the weakest survives

        • Diagram:

      • Symbiosis

        • Close and persistent long-term interaction between two species

          • Can be obligate (required for survival)

          • Or facultative (advantageous without being strictly necessary)

        • Symbiotic relationships can be beneficial to either one or both organisms in the partnership

          • Mutualism – both species benefit from the interaction

            • Example: how anemone protects clownfish and clownfish provides fecal matter for food

          • Commensalism – one species benefits, the other is unaffected

            • Example: barnacles are transported to plankton-rich waters by whales

          • Parasitism – one species benefits to the detriment of the other species

            • Example: ticks or fleas feed on the blood of their canine host

          • Diagram:

    • A: Local examples to illustrate the range of ways in which species can interact within a community

      • Mutualism

        • Ongoing interaction between two species whereby both species benefit from the interaction

        • Examples:

          • Honey bees gather nectar from flowers and distributes pollen between plants

            • This mediates the plant life cycle

          • Plover birds pick food morsels from between the jaws of crocodiles, cleaning their teeth in the process

          • Zooxanthellae photosynthesise within the protective environment of the polyp’s endodermis

            • This feeds the coral

      • Commensalism

        • Ongoing interaction between two species whereby one benefits and the other is unaffected

        • Examples:

          • Remora attaches to the underside of larger predatory fish and feed off the uneaten food scraps

          • Monarch butterflies can safely store poisonous chemical produced by milkweeds discourage predators from eating it

          • Decorator crabs remove small fragments of tissue from sea sponges and uses them as a source of camouflage

      • Parasitism

        • Ongoing interaction between two species whereby one species benefits at the other’s expense

        • Examples:

          • Ticks infest the skin and fur of host animals, feeding off the host and potentially causing disease

          • Leeches attach to the skin and drinks the blood of the host animal until fully engorged

          • Tongue-eating louses eat the tongue of a fish and may still later ingested food

    • A: The symbiotic relationship between Zooxanthellae and reef-building coral reef species

      • Reef-building coral will form a mutualistic symbiotic relationship with the photosynthetic unicellular algae Zooxanthellae

        • Coral are colonial organisms made up of individual polyps that are connected by a layer of living tissue

        • The algae lives within the cells of the coral’s endodermis (the innermost lining of the animal)

      • The coral provides the algae with a protective environment and source of inorganic compounds

        • Coral polyps secrete calcium carbonate to build a stony skeleton which encases the polyps and the Zooxanthellae

        • Coral polyps also recycle the waste products of the algae and supplies the Zooxanthellae with CO2

      • The Zooxanthellae in turn provides the coral polyps with a necessary source of nutrition

        • The alage supplies the coral with oxygen, glucose and other organic molecules that it produces via photosynthesis

        • The algae also helps the coral remove necessary waste products

      • Diagram of their mutualistic relationship:

      • Coral bleaching

        • It is the Zooxanthellae that gives the coral its vibrant pigmentation

        • When there is a large scale loss of Zooxanthellae from the coral due to environmental stress, bleaching of the coral occurs

          • This causes the coral to starve and die unless the Zooxanthellae are restored

        • Conditions which can cause coral bleaching include:

          • Changes in light availability

            • E.g. sedimentation may increase the opacity of the oceanic waters

          • Temperature increases

            • Water temperatures in excess of 30 degrees Celsius can irrevocably stress the Zooxanthellae

          • Ocean acidification

            • The buildup of CO2 concentrations in the ocean can lower pH and stress the Zooxanthellae

  • Keystone species

    • U: Community structure can be strongly affected by keystone species

      • Keystone species = a species that has a disproportionately large impact on the environment relative to its abundance

        • Name comes from the keystone in an arch → it fundamentally supports the whole structure and prevents it from collapsing

      • Keystone species may influence communities in a number of ways:

        • Predators can exert pressure on lower trophic levels to prevent them from monopolizing certain resources

        • Mutualists can support the life cycle of a variety of species within a community

          • E.g. pollinators and seed dispersal

        • Engineers can refashion the environment in a manner that promotes the survival of other species

          • Engineer = any species that creates, significantly modifies, maintains and/or destroys habitats

      • Numerous examples of keystone species within different communities, such as:

        • Sea stars (predators) prey on urchins and mussels, which prevents mussel overpopulation and coral reef destructions by urchins

        • Honey bees (mutualist) pollinate a wide variety of plant species, ensuring the continuation of the plant life cycle

        • Beavers (engineer) build dams that transform the environment in a manner that allows certain other species to survive

      • Keystone species are not the dominant species (most numerous) within a community nor do they have to be apex predators

      • Diagram of consequences of removal of keystone species:


C2 Communities and ecosystems

  • Trophic levels

    • U: Most species occupy different trophic levels in multiple food chains

      • Trophic level = the position an organism occupies within a feeding sequence

        • Producers occupy the first trophic level in a feeding sequence

        • Primary consumers feed on producers → second trophic level

        • Secondary consumers feed on primary → third trophic level

        • Further consumers occupy the subsequent levels

        • Diagram:

      • Food chain = shows linear feeding relationships between species in a community

        • Arrows represent the transfer of energy and matter

          • This transfer happens as one organism is eaten by the other

          • The arrows point in the direction of energy flow

        • Goes from the producer to the consumers

        • Most species’ feeding requirements means they will be in multiple food chains where they may occupy a different trophic level

          • As few species only feed on one other species

        • E.g.:

    • U: A food web shows all the possible food chains in a community

      • Links food chains together into more complex and interrelated feeding relationships

      • More representative of actual feeding pathways within an ecosystem because

        • Organisms can have more than one food source

        • Organisms can have more than one predator

        • Hence, organisms can also occupy more than one trophic level

      • Tip! When constructing food webs, position organisms at their highest trophic level (from left to right) as it keeps all arrows pointing in the same directions

      • Example:

    • EC: Ecological Pyramids

      • Ecological pyramids show the relative amounts of a specific component at the different trophic levels of an ecosystem

        • The three main types measure species numbers, biomasses and energy

      • Pyramid of Numbers

        • Shows the relative number of organisms at each stage of a food chain

        • Are usually shaped like pyramids → higher trophic levels cannot be sustained if there are more predators than prey

          • However, the shape may be distorted if a food source is disproportionately large in size/biomass compared to the feeder

            • E.g. a large number of caterpillars may feed on a single oak tree

      • Pyramid of Biomass

        • Shows the total mass of organisms at each stage of a food chain

        • Almost always upright in shape as biomass diminished along food chains as CO2 and waste is released

          • An exception to this rule is found in marine ecosystems where zooplankton have a larger total biomass than phytoplankton

            • Because phytoplankton replace their biomass at such a rapid rate and can therefore support a larger biomass of zooplankton

      • Pyramid of Energy

        • Shows the amount of energy trapped per area in a given time period at each stage in a food chain

        • Always upright in shape as energy is lost along food chains

          • Either used in respiration or lost as heat

        • Each level in the pyramid is roughly one tenth the size of the preceding level as energy transformations are ca. 10% efficient

      • Examples of pyramids:

  • Energy conversions

    • U: The percentage of ingested energy converted to biomass is dependent upon the respiration rate

      • Ecological production/productivity refers to the rate at which biomass is generated in an ecosystem

        • Usually expressed in units of mass per area per time

          • km m^-2 day^-1

      • Biomass = total dry weight of organic matter in organisms or ecosystems

        • The entirety of all biologically produced organic matter

          • E.g. proteins, carbohydrates, etc.

      • The percentage of energy that is converted into biomass is dependent upon a number of factors that lose energy

        • Energy is lost as inedible materials (bones, teeth, hair)

        • Energy is lost via excretion of undigested or unabsorbed materials

        • Energy is lost as heat from cellular respiration

          • The higher respiration rate results in more heat lost

      • Primary production = the production of chemical energy in organic compounds by producers

        • The main source of energy for primary production is sunlight

          • Sometimes chemosynthesis by lithotrophs

        • Can be categorized as one of two types:

          • Gross primary production (GPP) → the amount of chemical energy as biomass that a producer creates in a given length of time

          • Net primary production (NPP) → the amount of chemical energy that is not consumed by respiration

            • GPP - respiration

      • Secondary production ? the generation of biomass by heterotrophic organisms (consumers)

        • Biomass generation is driven by the transfer of organic compounds between trophic levels via feeding

        • Secondary production may also be categorized according to gross and net amounts of biomass

          • Gross = total

          • Net = usable

      • Diagram explaining how ingested energy converted to biomass depends on respiration:

  • Feed Conversion Ratio

    • A: Conversion ratio in sustainable food production practices

      • Feed Conversion Ratio (FCR) = mass of feed/mass of desired output

        • FCR measure the efficiency of an animal in converting food into a desired output

        • The lower the FCR the more efficient the method of food production

      • Low FCR is achieved by minimizing the potential losses of energy for the animal stock

        • Restricting animal movement will reduce energy lost to cell respiration

        • Optimizing feeding practices so that food is ingested and digested more effectively

        • Slaughtering animals at a young age

          • As older animals tend to grow more slowly and have a higher FCR

      • Theoretically the most efficient FCR is best as it lowers cost, however there are a number of potential ethical issues involved

        • Is it ethical to restrict animals and kill them early?

      • Diagram displaying the FCRs of different species:

    • EC: Sustainable food practices

      • Most of the protein found in Western human diets comes from farmed animal meet

        • E.g. cattle and poultry

        • These animals must be fed and housed → this creates an ecological footprint, the size of which depends on the FCR

      • Sustainable food practices provide food for the consumer while minimizing the environmental impact of the food source

        • The benefits of lowering food production costs must be balanced against ethical considerations regarding animal treatment

      • One sustainable food practice is the use of crickets as a protein source (as opposed to cattle)

        • Crickets have a substantially lower FCR → minimizes feed costs

        • They require significantly less water → minimizes water usage

        • They can reproduce much faster and require less space

        • Produce far less methane

        • For these reasons, certain health professionals are advocating the introduction of crickets into normal diets

          • The advantages of crickets as a food source is still debated though, so implementation is likely to be slow

  • Types of Ecosystems

    • U: In closed ecosystems energy but not matter is exchanged the with the surroundings

      • Ecosystems can either be open or closed, depending on whether matter moves into and out of the system

        • A closed ecosystems exchanges energy with its surrounding, but not matters

          • It is self-contained

          • E.g. a mesocosm

        • An open ecosystem changes both energy and matter with surrounding environments

          • A natural ecosystem such as a forest

    • U: The type of stable ecosystem that will emerge in an area is predictable based on climate

      • Ecosystem = the interaction of living and non-living things within an area

        • A community and its abiotic environment

      • Particular types of stable ecosystems will emerge in a given geographical area according to its climate conditions

      • Biome = a geographical area that has a particular climate and sustains a specific community of plants and animals

        • Basically a specific type of natural ecosystem, created from its climate and environment

        • The main factors affecting the distribution of biomes is temperature and rainfall

        • These factors vary according to latitude, longitude, altitude, proximity to ocean, etc.

      • Many different biomes are found across continents, including:

        • Tropical rainforests

          • Hot and humid environments

          • Near the equator

          • Dense vegetation

          • High biodiversity

        • Taiga

          • Coniferous forests

          • Near the poles

          • Have cold temperatures

          • Little precipitation

            • Moisture trapped as snow or ice

        • Deserts

          • Dry and arid environments

          • Display extreme temperature conditions (hot and cold)

        • Picture examples:

    • EC: Types of Biomes

      • Biomes are geographical area that have a particular climate and sustain a specific community of flora and fauna

        • The total area where all living things are found is called the biosphere

          • The totality of biomes

        • Certain biomes are named and recognized according to their specific characteristics

      • Tropical rainforest

        • Hot climate → 25-30 degrees Celsius

        • Very high levels of precipitation → > 250 cm per year

        • Large diversity in species

        • Vegetation includes epiphytes, tall trees and undergrowth

          • Epiphytes: organisms that grow on the surface of plants

      • Temperate forest

        • Also called deciduous forests

        • Moderate temperatures and clear seasonal changes

        • Growing period of around 200 days during 4-6 frost free months

      • Boreal forest (Taiga)

        • Coniferous forests

        • Cold and icy → 0-15 degrees Celsius

        • Only small amount of precipitation

        • Coniferous trees are densely packed and there is little variation in species

      • Tropical Grassland (Savannah)

        • Warm → 20-30 degrees Celsius

        • Medium amounts of moisture, although seasonal droughts are common

        • Trees are intermittent and widely spaced

        • Occasional fires prevents thicker vegetation from encroaching

      • Temperate Grassland

        • Moderate temperatures and amounts of rainfall

        • Trees and shrubs are largely absent

        • Grass is the dominant vegetation

      • Mediterranean (Chaparral)

        • Moderate temperatures → 15-25 degrees Celsius

        • Rainy winters and dry summers

        • Vegetation consists of dry woody shrubs that are quick to grow

      • Desert

        • Extreme temperature conditions → > 30 degrees Celsius during the day, < 0 degrees at night)

        • Very low precipitation

          • < 30 cm per year

        • Dominant plant species are xerophytes which are adapted for water conservation

      • Tundra

        • Freezing temperatures → < 0 degrees Celsius

        • Very little precipitation

        • Vegetation is low growing

          • E.g. mosses

        • Perennial plants may grow during the summer

      • Mountain ranges

        • Found all over the world at high attitudes → > 10 000 feet

        • Temperatures are typically low due to the altitude

        • Weather conditions may change rapidly

      • Distribution of biomes in the biosphere:

  • Ecosystem Analysis

    • S: Analysis of a climograph showing the relationship between temperature, rainfall and the type of ecosystem.

      • Climograph = graphical representation of basic climate parameters at a given geographical location

        • Shows a combination of monthly average temperature and precipitation at a certain location

          • Precipitation = rainfall

        • Provides a quick overview of the climate of a region and can be used to identify seasonal patterns and changes

        • Examples:

      • Whittaker’s climograph

        • In 1975, ecologist Robert Whittaker developed a holistic climograph that allowed for the classifying of 9 distinct biomes

        • Biomes distinguished according to their average yearly temperatures and precipitation

          • Deserts typically have high average temperatures but low precipitation

            • Hot and dry

          • Rainforests typically have high average temperatures and high precipitation

            • Hot and wet

          • Taigas typically have low average temperatures and reasonably low precipitation

            • Cold and icy

    • S: Comparison of pyramids of energy from different ecosystems

      • Pyramids of energy (PoE) = diagram of the flow of energy between trophic levels

        • Measures in units of energy per area per time

        • Standard units are kJ m^-2 and y^-1

        • Triangular and never inverted as 90% of energy is lost between trophic levels

      • PoEs differ between ecosystems due to the effect of climate on primary productivity

        • Warmer temperatures speed up enzyme reactions required for photosynthesis

        • High precipitation also increases photosynthesis as the photolysis of water is essential for non-cyclic photophosphorylation

        • Higher (optimal) rate of photosynthesis = more energy transferred

      • Tropical rainforests have a high NPP whereas deserts have a low NPP

        • Higher NPP = more energy supplies to consumers hence more trophic levels can exist

        • Tropical rainforests PoE has a wider base and more levels than a desert

      • Types of species within a particular biome may also affect how efficiently energy is transferred between trophic levels

        • Example: Homeotherms will use more energy maintaining a stable body temperature compared to poikilotherms

      • Diagram:

    • S: Construction of Gersmehl diagrams to show the interrelationships between nutrient stores and flows between taiga, desert and tropical rainforest

      • Gersmehl diagrams show the differences in nutrient flow and storage between different types of ecosystems

      • Nutrients are stored in one of three nutrient sinks

        • Biomass → the total mass of living organisms in a given area

          • Mainly plant tissue

        • Litter → any organic matter in and on the soil

          • Includes humus and leaf litter (defoliation)

        • Soil → the top layer of the earth that is composed of disintegrated rock particles

      • Nutrients can be transferred between nutrient sinks and may also be cycled via environmental inputs and outputs

        • Transfer from biomass to litter → fallout

        • Transfer from litter to soil → decay

        • Transfer from soil to biomass → uptake

        • Litter can additional gain nutrients via precipitation and lose nutrients in surface runoff

        • Soil can gain nutrients from the erosion of rocks via weathering and lose via leaching

      • Standard: Gersmehl diagram

      • Comparing Gersmehl diagrams

        • The inter-relationships between nutrient stores and flows will be affected by climatic factors

          • Such as temperature and rainfall

          • Therefore Gersmehl diagrams will differ between biomes

        • Tropical rainforest

          • Most nutrients are stored as biomass

            • Litter is rapidly decomposed and vast roots quickly draw nutrients from soil

          • There is a fast rate of transfer between stores

            • Hot and wet conditions promote precipitation, runoff, weathering and leaching

        • Taiga

          • Most nutrients are stored as litter

            • Low temperatures slow decomposition which delays nutrient transfer to soil and biomass

          • There is little nutrient gain from precipitation or weathering due to low levels of precipitation

            • Cold temperatures produce snow instead

          • There is little nutrient loss via leaching due to low precipitation

            • But surface runoffs may be high at certain times if the snow melts

        • Desert

          • Most nutrients are stored in the soil

            • Few plants exist to store nutrients as biomass nor to produce litter

          • There is little nutrient gain from precipitation and little nutrient loss via runoff → both due to the dry climate

          • The amount of weathering and leaching is insignificant

  • Primary Succession

    • S: Analysis of data showing primary succession

      • Ecological succession = the process by which a sequence of increasingly complex communities develop over time

        • Climax community = when succession has ended and the community has all of its characteristics

      • Primary succession = when communities develop on entirely new land without any established soil

        • May occur at river deltas, glaciers, sand dunes or on exposed rock

        • First organisms to colonize the region are called pioneer species

          • Typically consist of lichen or moss

            • When they die, they decompose which creates the first organic soil capable of sustaining plant growth

      • As plant species colonize a new area, the litter produced by their growth and decomposing remains will cause changes

        • Soil depth will increase because plants add humus to the soil

        • Soil pH will become altered by the additions

        • Soil mineral content will increase and rocks will begin to be broken down by the action of roots

        • The soil will become aerated and water retention increases

          • Drainage is reduced

        • These changes will allow for growth of larger plants, which will reduce erosion through the binding action of their roots

          • Not all plant species will thrive in unison → larger plants will eventually outcompete smaller shade intolerant plants

      • Overview of primary succession:

      • Primary succession data

        • A primary succession sequence can be identified according to the distribution of plants

          • This applies for any location where a new community may emerge from uninhabitable land

          • The regions closer to the site of development will be in the earlier chronological stages of succession

            • E.g. Glacial retreat exposes bare rock for colonization, so regions further from the glacier have had more time to develop life

        • Diagram example:

  • Ecological Disturbance

    • U: Disturbance influxes the structure and rate of change within ecosystems

      • Environmental disturbances may cause fluctuations to the structure and rate of change within ecosystems

        • Such disturbances may be natural or artificial (human-induced)

          • E.g. bushfire, earthquake

      • Natural disturbance

        • A natural environmental disturbance may give rise to secondary succession

          • Secondary succession = one ecosystem is replaced by another

          • Occurs when succession starts on existing soil following the upheaval (change) of a pre-existing ecosystem

            • This upheaval results in the removal of existing biota and allows a new ecosystem to develop on the site of the old

          • Because the soil is already developed during secondary succession, dominance is usually achieved by the fastest growing plants

        • The progression of secondary succession can be summarized as follows:

          • An environmental disturbance destroys the pre-existing climax community

          • Grasses and herbaceous plants are the first to grow back as the soil is already present

            • Soil = no pioneer species required

          • Fast growing trees will develop to their fullest while shade tolerant trees will develop in the understory

            • Eventually the fast-growing trees may be overtaken by larger, slower-growing trees as the ecosystem reverts to its prior state

        • Diagram:

    • A: Consideration of one example of how humans interfere in nutrient cycling

      • Deforestation is the permanent destruction of a forest via the removal or clearance of trees

        • Human-induced degradation of forests

        • Driven primarily by a need for timber and cleared land for agricultural purposes

      • Deforestation disturbs the normal nutrient cycling within the region in a number of ways

        • Less trees means there is less evapotranspiration  meaning there is less moisture/precipitation in the air

        • Fewer trees means less litter

          • Due to less defoliation (leaves falling off a plant)

          • This reduces the production of humus, so there will be less nutrients in the soil

        • Rapid loss of nutrients from leaching

        • Less chemical weathering of rock, due to less water in soil

        • The soil will become more acidic and release iron and aluminum to form an infertile ferralitic soil

          • Ferralitic soil = nutrient poor

        • The soil layer will become increasingly thin and eroded

        • The infertile soil will prevent vegetative growth, reducing biodiversity and nutrient cycling

        • Logging operations may also alter the distribution of plant species by removing the canopy and increasing light exposure

          • Canopy = the upper layer of habitat zone, usually formed by mature tree crowns

        • Removal of the canopy also results in an increases loss of nutrients from the soil via runoff

      • The effect of deforestation on nutrient cycling:

    • S: Investigation into the effect of an environmental disturbance on an ecosystem

      • Environmental disturbances are caused by natural or artificial disruptions to a normal ecosystem

        • Examples:

          • Fire breaks in bush lands or regions damaged by bushfires

          • Outer boundaries of population settlements or regions bordering roads

          • Dams and artificial rivers and creeks

            • E.g. irrigation sites

      • The effect of an environmental disturbance on an ecosystem can be measured in a number of ways

        • Population density

          • Using the Lincoln index via the capture-mark-recapture technique

        • Species diversity and richness

          • Using the Simpson’s reciprocal index

        • The presence and distribution of indicator species

          • To measure levels of pollution

        • Canopy coverage and relative light intensity

          • Could measure with a lux meter

        • Biomass

          • Via the average width of tree stems at a specified height

        • Edaphic factors such as soil erosion (via depth), water retention (via drainage), pH and nutrient content

      • Measurements taken from a disturbed area need to be compared against measurements taken from an undisturbed control area

        • This enables the investigator to statistically calculate both the effect and magnitude of the environmental disturbance

C3 Impacts of humans on ecosystems

  • Invasive species

    • U: Introduced alien species can escape into local ecosystems and become invasive.

      • The species within any given ecosystem can be broadly described as either endemic or alien

        • Endemic = native species to a defined geographic region

          • E.g. koalas in Australia

        • Alien = species that have been transferred from their natural habitat into a new environment

        • If an introduced alien species has detrimental effects on the pre-existing food chains, it is classified as invasive

          • Invasive species typically threaten the biodiversity of the ecosystem they are introduced into by displacing native species

      • Example: invasive species in Australia → cane toad, European rabbit, red fox, camel, feral cat

    • U: Competitive exclusion and the absence of predators can lead to reduction in the number of endemic species when alien species become invasive

      • Competitive exclusion principle = two species cannot occupy identical niches within a community

        • One species will have a competitive advantage and survive at the expense of the other

        • Evolution via natural selection results in adaptations that function to minimize direct competition between species in nature

      • Invasive species possess several advantages that allow them to displace endemic species via competitive exclusion:

        • Typically possess a large fundamental niche

          • = can occupy a wider area than endemic species

        • Commonly lack a predator capable of limiting their survival

          • As they are introduced into a foreign environment

        • May possess certain features that are suited to the new environment

          • E.g. more aggression, better foraging, etc.

      • Diagram:

    • A: Study of the introduction of cane toads in Australia and another local example of an introduced alien species

      • Case study one: Cane toads (Bufo marinus)

        • Cane toad was introduced in Australia in 1935 by humans to control the spread of the sugar cane beetle that was damaging crops

          • Failed to limit the beetle population → beetles live at the tops of sugar canes where the toads could not reach

        • The cane toad has reproduced rapidly and is now classified as an invasive species that endangers native wildlife

          • It has a wide diet → depleted the prey population for native insectivores

          • Secretes a toxic chemical from its skin that has poisoned native predators

          • Carries diseases that have been transmitted to native species of frogs and fish

        • The cane toad has spread across the northern fringes of Australia

          • Now numbers in the millions

        • Proven difficult to eradicate as there are no natural predators in Australia to reduce its population size

      • Case study 2: Wild rabbits

        • European rabbits were introduced in Australia with the arrival of the First Fleet

          • Ostensibly for recreational hunting

        • Australian conditions promoted a population explosion

          • Mild winters allowed rabbits to breed the entire year

          • Fastest spread ever recorded for any mammal anywhere in the world

        • Rabbits have had a devastating effect on native Australian ecology

          • Killed native plant species by ringbarking

            • Ringbarking = removing a strip of bark that prevents future growth beyond that point

          • Loss of native plants has left the topsoil exposed and vulnerable to extensive erosion from wind and rain

        • Various strategies have been employed to control rabbit populations with varying effect

          • ‘Rabbit-proof’ fences were built to try and contain rabbit populations

          • Methods of hunting and poisoning have been utilized

            • Also efforts to destroy rabbit warrens (burrows)

          • Rabbit-borne diseases have been released with moderate success

            • E.g. myxomatosis virus

  • Biological control

    • S: Evaluation of eradication programmes and biological control as measures to reduce impact of alien species.

      • Controlling the population of invasive species once they have become established is difficult and typically very expensive

        • Three main methods: physical, chemical, biological

      • Physical control

        • Involves removal or restriction by manual or mechanical measures

        • May include:

          • Installation of barriers and fences

          • Removal of habitat by excavation or trimming

          • Population numbers may be reduced by hunting, trapping and culling

            • Although this is labor intensive

        • Physical methods to contain invasive species are not usually species specific and can also impede endemic wildlife

      • Chemical control

        • Involves the use of chemical agents to limit population numbers and spread

          • Chemical agents = poisons and toxins

        • Chemical agents may include:

          • Herbicides for plants

          • Pesticides for insects

          • Other compounds such as rat poison

        • Chemical agents may have moderate specificity, but can also detrimentally affect local wildlife + are costly

        • The effect may become more pronounced in high trophic levels due to biomagnification

      • Biological control

        • Involves using living organisms or viruses to control an invasive species

        • Biological control may eat the invasive species or cause it to become diseased

        • Biological agents mut be carefully assessed before release to ensure they do not become invasive themselves

        • Examples:

          • The Vedalia beetle feeds on citrus plant invertebrates

          • The Myxoma virus that affects rabbits

        • Biological control agents must be monitored for unintended side effects

          • E.g. development of immunity in invasive species

        • Biocontrol case study: The Vedalia Beetle (ladybug)

          • The cottony cushion scale is an invertebrate pest from Australia that was accidentally released in California

            • Spread and fed on citrus plants such as orange trees

            • Devastated the Californian citrus industry

          • The Vedalia beetle is a predatory insect from Australia that was introduced into California as a means of biological control

            • It worked to limit the numbers of the cottony cushion scale

            • Minimized the economic impact on the citrus industry

          • Diagram:

  • Biomagnification

    • U: Pollutants become concentrated in the tissues of organisms at higher trophic levels by biomagnification

      • Biomagnification = the process in which chemical substances become more concentrated at each trophic level

        • Bioaccumulation = the build up of a chemical substance in the tissues of a single organism

      • Biomagnification occurs because organisms at higher trophic levels must consume more biomasse to meet feeding requirements

        • Energy transformations are only ca. 10% efficient, so higher order consumers must eat more to meet energy demands

        • This means higher order consumers will experience increased contamination from a chemical substance

      • Diagram of effect:

    • S: Analysis of data illustrating the causes and consequences of biomagnification

      • Dichloro Diphenyl Trichloroethane (DDT) is an example of a chemical substance which is biomagnified

        • DDT = a chemical pesticide that is sprayed on crops to eliminate mosquito larvae and washed into waterways at low concentrations

          • Taken up by algae and passed on to primary consumers

        • DDT is fat soluble and selectively retained within the tissues of an organism instead of being excreted

      • At each subsequent trophic level DDT is passed on through the algae, the concentration of DDT stored in the body increases

        • Increased trophic level = increased food uptake = increased DDT

        • Very high levels of DDT were discovered in birds that preyed on fish

      • Birds exposed to high levels of DDT through feeding on fish that fed on algae were found to produced thinner shells as a consequence

        • This decreased survival rates of fledglings

      • Diagrams we may encounter for such data analysis

    • A: Discussion of the trade-off between control of the malarial parasite and DDT pollution

      • DDT was widely employed as a chemical pesticide in the 1940s and 1950s to control insect-borne diseases like malaria

        • Agricultural insecticide

        • Led to complete eradication of malaria in many high socioeconomic world regions

        • Less effective in low socioeconomic tropical regions where eradication programs could not be sustained

      • Eradication programs were curtailed by the World Health Organization (WHO) in 1969

        • For safety and environmental concerns

        • DDT can bioaccumulate to toxic levels and poses a significant threat to high trophic levels due to biomagnification

        • Current interventions for malaria now focus on non-spraying strategies

          • E.g. the use of bednets soaked in insecticides

      • Malaria vs. DDT

        • Arguments for DDT spraying:

          • DDT spraying is an affordable and effective means of killing mosquitos that carry disease

            • E.g. malaria, dengue fever, etc.

          • Where the use of DDT has been discontinued, the incidence of malaria and associated deaths have increased

          • Health costs associated with the treatment of malaria are reduced when DDT spraying is implemented

          • Alternative strategies are not as cost-effective or successful

        • Arguments against DDT spraying

          • It is associated with adverse health effects in humans

            • E.g. cancer, birth defects, reduced fertility, etc.

          • Persists in the environment for long periods of time

            • More than 15 years

          • DDT is biomagnified in higher order consumers which has adverse consequences on ecosystems

  • Plastic pollution

    • U: Macroplastic and microplastic debris has accumulated in marine environments

      • Plastic = a type of synthetic polymer

        • Found in certain types of clothes, bottles, bags, food wrappings, containers, etc.

        • Most plastics are not biodegradable → persist in the environment for many centuries

      • Macroplastic = large visible plastic debris

        • 1 mm

        • Macroplastic debris can be degraded and broken down into microplastic debris by UV radiation and the action of waves

      • Microplastic = smaller plastic debris

        • < 1 mm

      • Ocean currents will concentrate plastic debris in large oceanic convergence zones called gyres

      • Plastic debris in the ocean will leach chemical into the water + absorb toxic contaminants called persistent organic pollutants

        • Microplastics will absorb more persistent organic pollutants (POPs) due to their smaller size

          • More surface area available

      • Both macro- and microplastic debris is ingested by marine animals, which mistake the debris for food

        • This leads to the bioaccumulation and biomagnification of POPs within marine animals

        • It may also damage the stomach of animals or cause them to stop feeding

          • By taking up space in the digestive tract

      • Diagram:

    • A: Case study of the impact of plastic debris on Laysan albatrosses and one other named example

      • Case Study 1: Laysan Albatross

        • Nests on islands found in the North Pacific gyre where large amounts of plastic debris is found

        • Feeds by skimming the ocean surface with their beak → this causes them to ingest large quantities of plastic too

        • Adults can regurgitate (throw up) the plastic they have swallowed, but chicks are unable to

          • Instead, it fills up their stomachs

          • The mortality rate in albatross chicks is very high

            • Estimated 40% die before fledgling

      • Case Study 2: Sea turtles

        • Will commonly mistake plastic bags for jellyfish

          • Jellyfish = one of their primary food sources

        • Ingestion of the plastic can be fatal

          • It can become lodged in the esophagus and cause future feeding problems

        • Plastic debris can also become wrapped around the turtle → restricting movement and developmental growth

        • It is estimated plastic pollution harms ca. 100 000 sea turtles and other marine animals each year

  • EC: Cholorfluorocarbons

    • Chlorofluorocarbons (CFCs) = chemicals widely used as refrigerants, propellants and solvents in the past

      • Contribute to ozone depletion in the upper atmosphere

        • Therefore their manufacture and use is being phased out

    • Ozone layer = stratospheric region composed of ozone (triatomic oxygen)

    • CFCs are broken down by UV radiation to release chloride ions that react with ozone

      • The ions breaks the ozone down into oxygen

      • One chloride ions can destroy 100 000 ozone molecules

      • Diagram of the process:

    • Ozone in the stratosphere absorbs UV radiation, however there is a limit to how much radiation ozone can absorb

      • Regions where the ozone layer is thinner will experience higher terrestrial UV levels

        • Thinning caused by for example CFCs

      • UV light penetrates tissues and damages DNA → this causes gene mutations and cancer

      • Can also damage the ability of plants to carry out photosynthesis + kill phytoplankton → this reduces productivity

  • EC: Earth spheres

    • The Earth is made up of four distinct and connected spheres

      • Lithosphere, hydrosphere, atmosphere, biosphere

    • Lithosphere = all the rocks, minerals and molten magma found on or in the Earth

      • Also called geosphere

    • Hydrosphere = all the water on Earth

      • Liquid water such as oceans and rivers

      • Precipitation and vapor

    • Atmosphere = the layer of gasses surrounding the Earth

      • Divided into further sections

      • E.g. stratosphere

    • Biosphere = all the living organisms on the planet

      • Including plants, animals, bacteria, etc.

    • As the four spheres are interconnected, human impact on one sphere will potentially affect other spheres

      • Examples:

        • The release of plastic pollution into the hydrosphere (oceans) will impact the biosphere (marine life)

        • The production and release of CFCs into the atmosphere will impact how much UV radiation the biosphere is subjected to

C4 Conservation of biodiversity

  • Indicator species

    • U: An indicator species is an organism used to assess a specific environmental condition

      • Indicator species are sensitive to specific environmental conditions → limited range of tolerance

        • Reduction or growth of their population therefore indicates changes in the environment

          • Useful means of monitoring change

      • Examples of indicator species and their environmental conditions

        • Lichen and mosses are susceptible to air-borne pollutants dissolved in water

          • E.g. sulfur dioxide

        • Tubifex worms are sensitive to concentrations of heavy metals

        • Mayfly larva and certain aquatic invertebrates are sensitive to dissolved oxygen levels in water

      • Diagram:

    • U: Relative numbers of an indicator species can be used to calculate the value of a biotic index

      • Biotic indices compare the relative frequency of indicator species

          • Provide an overall assessment of environmental health

      • Calculating a biotic index involves multiplying the population size of each indicator species by its pollution tolerance rating

      • The following conclusions can be drawn from a biotic index:

        • A high biotic index = the presence of many pollution-sensitive organisms

          • Denotes an unpolluted environment

        • A low biotic index = the absence of pollution-sensitive organisms + abundance of pollution-tolerant organisms

          • Indicates a polluted environment

        • A change in the biotic index over time marks a change in the environmental conditions within a given ecosystem

      • Pollution tolerance rating

      • Environmental pollution levels

  • Biodiversity

    • U: Richness and evenness are components of biodiversity

      • Biodiversity = the variety and variability of all living organisms within a given ecological area

        • Can be used to refer to the number of species, their genetic diversity or habitat variety

          • Habitat variety = ecological variations

      • 2 main components contribute to biodiversity

        • Species richness → the number of different species present in an area

          • More species = greater richness

        • Species evenness → the relative abundance of the different species in an area

          • Similar abundance = more evenness

        • Richness vs. evenness

    • S: Analysis of the biodiversity of two local communities using Simpson’s reciprocal index of diversity

      • Simpson’s reciprocal index can be used to measure the relative biodiversity of a given community

        • It takes into account both the number of species present (richness) and the number of individuals per species (evenness)

        • A higher index value is indicative of a greater degree of biodiversity within the community

      • Simpson’s reciprocal index can be used to compare communities to identify intrinsic values

        • Intrinsic value = something we value for itself, not its consequences

        • A high index value = a stable site with many different niches and low competition

          • High richness and evenness

        • A low index value = a site with few potential niches where only a few species dominate

          • Low richness and evenness

        • The index value may change in response to ecological disturbance

          • E.g. human intervention, natural disasters, etc.

  • Biogeographic factors

    • U: Biogeographic factors affect species diversity

      • Biodiversity within a given ecosystem will be affected by a number of biogeographic factors

        • Larger habitats tend to promote biodiversity better than smaller ones

          • More available niches = less competition

        • Ecology at the edges of ecosystem is different from central areas

          • E.g. more sunlight, more wind, etc.

          • This is known as the edge effect, whereby species distribution is influenced by divergent environmental conditions

          • Edges tend to have greater biodiversity as different habitats with different abiotic factors exist in close physical proximity

          • However, edges tend to have more competition than central regions → may restrict survival prospects of certain species

        • Habitat corridors between parts of a fragmented habitat can connect disparate regions to improve genetic diversity

      • These principles of biogeographic factors are routinely applied when setting aside land as a nature reserve to improve the conservation of biodiversity

    • A: Analysis of the impact of biogeographic factors on diversity limited to island size and edge effect

      • Island size

        • The biodiversity of an island is typically proportionate to island size

          • Larger island = greater biodiversity

            • Larger islands support a greater range of habitats

              • Hence more available niches for species to occupy → more survival + diversity

            • Larger islands can sustain higher population numbers for each species

              • Increases species evenness

            • Larger islands have greater productivity at each trophic level

              • Leads to longer + more stable food chains

          • Smaller island = less biodiversity

          • Diagram depicting the relationship between island size and biodiversity

      • Edge effect

        • The diversity of species within a given environment changes with proximity to the ecosystem’s boundaries

        • Biodiversity may be greater at the border between two ecosystems as different abiotic factors favor certain species

        • However, certain species may not be able thrive under these conditions

          • Instead must occupy more central regions

        • The effect of edges on biodiversity will be greatly influenced by the particular conditions caused by the ecological disturbance

        • Diagram:

  • Conservation

    • U: In situ conservation may require active management of nature reserves or national parks

      • Conservation = the careful preservation, protection and maintenance of natural resources

        • Such as trees, water, wildlife, etc.

        • Can be either in situ or ex situ

          • On site or off site

      • In situ conservation is the preservation of plant and animal species within their natural habitat

        • Typically involves the designation of protected areas of land as either nature reserves or national parks

      • These areas may require active management to ensure that an appropriate and sustainable ecological balance is maintained

        • Ecological monitoring of a species may be required to ensure viable population levels are maintained

        • Interventions may be required to prevent habitat degradation or competition from invasive species

        • Legislation may be necessary to ensure adequate funding for policing and education

      • In situ conservation offers several advantages when protecting endangered species from extinction

        • It allows species to live in the environment to which they are adapted

          • + to occupy their natural position in the food chain

        • It maintains the animal’s normal behavior

          • Offspring usually acquire skills from parents and peers around them

        • Retaining the natural habitat prevents its eventual loss + ensures it remains available for other endangered species

          • Such areas provide a place to return animals from breeding programs as they provide realistic conditions for reintegration

        • Reserves in different areas can share information and provide a place for scientific study and developing public awareness

    • U: Ex situ conservation is the preservation of species outside their natural habitats

      • Ex situ conservation involves the preservation of plant and animal species outside their natural habitats

        • May typically be required for critically endangered species when urgent intervention is necessary

      • There are several advantages associated with ex situ conservation

        • It allows for greater control of essential conditions

          • E.g. climate control, dietary intake, veterinary care, etc.

        • It can improve the chances of successful breeding by utilizing artificial methods

          • E.g. embryo transfer, IVF, etc.

      • Ex situ conservation is also associated with several disadvantages

        • They do not prevent the potential destruction of their natural habitats

        • Species raised in captivity are less likely to be successfully reintroduced into the wild

          • Loss of autonomous survival

        • Increases inbreeding by restricting the gene pool → restricts the evolution of the species

      • Some examples of ex situ conservation measures employed around the world:

        • Captive breeding → animals are raised and bred in containment to ensure survival prospects

          • E.g. zoos

        • Botanical gardens → areas devoted to the collection, cultivation and display of a wide variety of plant species

        • Seed banks → secure sites that store and catalog seeds in order to preserve plant genetic diversity

    • A: Case study of the captive breeding and reintroduction of an endangered animal species

      • Case Study 1: Indian rhinoceros

        • An endangered species with only ca. 3500 rhinoceros living in the wild

          • This number was as low as 2000 in the early 1990s → increased due to successful captive breeding programs

        • In addition to habitat loss, the Indian rhinoceros is threatened by poachers

          • Their horn is considered potent in some cultures

        • It is listed as vulnerable on the IUCN red list + the majority of the species is situated in Indian protected areas (reserves)

      • Case Study 2: Mountain Chicken Frog

        • A species of frog native to the Carribean islands of Dominica and Montserrat

        • The population of this frog has declined i81% in the last 10 years

          • Due to the fungal disease chytridiomycosis

          • Also threatened by human consumption → local delicacy said to taste like chicken

        • Considered critically endangered with fewer than 8000 individuals existing in the wild

        • The frog has been artificially bred in laboratories in England prior to being reintroduced into the wild

  • EC: Endangered species

    • Endangered species = categorized by the International Union for Conservation of Nature (IUCN)  as likely to become extinct

      • Conservation status = likelihood of becoming extinct

    • Currently 3000 endangered species according to IUCN

    • Factors for determining a species’ conservation status

      • Rate of decline

      • Population size

      • Area of geographic distribution

      • Degree of population fragmentation

    • IUCN has a red list which classifies species into nine groups

      • Extinct (EX)

        • No surviving individuals

        • E.g. tasmanian tiger

      • Extinct in Wild (EW)

        • Captive individuals survive, but there is no wild population

        • E.g. barbary lion

      • Critically endangered (CR)

        • Faces an extremely high risk of extinction in the near future

        • E.g. red wolf

      • Endangered (EN)

        • Faces a high risk of extinction in the near future

        • E.g. snow leopard

      • Vulnerable (VU)

        • Faces a high risk of endangerment in the medium term

        • E.g. Indian rhinoceros

      • Near Threatened (NT)

        • May be considered threatened in the near future

        • E.g. tiger shark

      • Least Concern (LC)

        • No immediate threat to species survival

        • E.g. giraffe

      • All other organisms may be listed as Data Deficient (DD) or Not Evaluated (NE)

  • EC: Extinction

    • Extinction = the complete cessation of a species or higher taxon level, reducing biodiversity

    • Can occur gradually as one population of organisms evolves into something else

      • Phyletic extinction

    • Can be sudden without any identifiable descendents → cease to exits

      • Abrupt extinction

    • Can be very difficult to determine the moment of extinction → most categorizations are usually done retrospectively

      • Occasionally species thought to be extinct can be rediscovered after a period of time

        • E.g. lazarus taxa

    • 99% of all species that ever lived on Earth are considered to be extinct

      • Estimated to amount over 5 billion species

      • Natural catastrophes may cause mass extinction events

        • Large scale loss of species in relatively short period

    • Case Study: Tasmanian Tiger

      • Thylacinus cynocephalus

      • Became extinct after the arrival of European settlers to Australia

      • Tasmanian tigers would feed on introduced sheep → therefore hunted and poached

      • The loss of habitat due to human development + lack of successful breeding programs = population numbers dwindling

        • Aboriginal rock paintings suggest the Tasmanian tiger once lived on the mainland, but died out from predation and/or competition

      • Last Tasmanian tiger died in captivity in 1936

        • Declared extinct by international standards in 1986

CC

Option C: Ecology and Conservation

C1 Species and Communities

  • Survival factors

    • U: The distribution of species is affected by limiting factors

      • Limiting factor = a component of an ecosystem which limits the distribution or numbers of a population

        • Defines optimal survival conditions according to its effect on a species when in deficiency or excess

        • Can be biotic or abiotic

          • Biotic factors include interactions between organisms

            • Either intraspecific (within a species)

            • Or interspecific (between species)

          • Abiotic factors include environmental conditions

            • E.g. light, temperature, salinity (concentration of salts in water or soils), rainfall, wind velocity, soil pH, etc.

        • Typical abiotic limiting factors for plants within a community

          • Temperature

            • Plants can only survive within a narrow range of temperatures to which they are adapted

            • High temperatures will increase the rate of water loss by evaporation and may also denature metabolic enzymes

            • Low temps may cause plant sap to freeze

              • The expansion of frozen water in xylem can cause trunks to split

            • Certain species of woody plants synthesis antifreeze protein to prevent crystal formation in frozen cells

              • E.g. maple trees

          • Water availability

            • Water is needed for photosynthetic processes + maintaining cell turgor

            • Xerophytes are plant species that are adapted to survive in dry and arid environments such as deserts

              • E.g. cacti

            • Hydrophytes are plant species that are adapted to survive in frequently waterlogged soils

              • E.g. rice

          • Light availability

            • Light is essential to the process of photosynthesis whereby plants produce organic molecules

            • Low-growing plants will typically possess darker leaves in order to optimize their light conversion

              • Dark = more chlorophyll

            • Certain seaweeds have pigments adapted to absorbing blue wavelengths

              • E.g. kelp

              • Red does not easily penetrate water

        • Typical abiotic limiting factors for animal within a community

          • Temperature

            • Animal survival typically relies on narrow temperature range

              • Few animals can survive temperature extremes

            • Poikilotherms cannot maintain thermal homeostasis and must occupy environments according to temperature needs

            • Homeotherms can regulate their own internal body temperatures and hence can occupy a wider range of habitats

            • Body size will play a fundamental role in determining an animal’s capacity to retain heat

              • The surface area to volume ratio

          • Territory

            • Territorial boundaries may determine an animal's capacity to attract mates, rear young, forage for food and avoid predators

            • Territories may be temporary (e.g. migration, breeding sites) or alternatively may be permanent

            • In certain species, juveniles may have different environmental requirements to adults

              • E.g. tadpoles and frogs

            • Establishment of territories can lead to intraspecific and interspecific competition

          • Food availability

            • Availability of food supply is a critical determinant in influencing population size and distribution

            • Animals may require a particular plant or animal species as a food source

              • This limits their spread to certain regions

              • Seasonal or geographic variations may directly affect food availability within a population

                • E.g. seasonal migrations

      • Law of Tolerance

        • The Law of Tolerance was proposed by Victor Ernest Shelford (American zoologist) in 1911

        • States that populations have optimal survival conditions within critical minimal and maximal thresholds

          • When populations are exposed to the extremes of a particular limiting factor (too much or too little), the survival rates drop

        • The distribution of species in response to a limiting factor is a bell-shaped curve with 3 distinct regions

          • The optimal zone

            • Central portion of a curve which has conditions that favor maximal reproductive success and survivability

          • Zones of stress

            • Regions flanking the optimal zone where organisms can survive but with reduced reproductive success

          • Zones of intolerance

            • Outermost regions in which organisms cannot survive

            • Represents the extremes of the limiting factor

          • Example:

    • A: Distribution of one animal and one plant species to illustrate limits to tolerance and zones of stress

      • Plant example

        • Plant growth varies greatly in response to salinity levels (concentrations of salt within the soil)

        • Plants that are not salt tolerant are called glycophytes

          • They are easily damaged by high salinity

          • Most plant species are considered glycophytes

        • Plants that are salt tolerant are called halophytes

          • They may become stressed in freshwater environments

          • Less than 2% of all plants are halophytes

        • Cultivation of land for agriculture causes the water table to rise and concentrates salt at the roots

          • Such cultivation includes irrigation and grazing

          • This makes it harder for glycophytes to extract water from the soil and the uptake of salt can be toxic to the plant

        • Understanding salt tolerance for different plant species is therefore critically important to effective crop farming

      • Animal example

        • Coral species form connected reefs → these are greatly impacted by changes in oceanic temperature

        • Coral polyps receive nutrition from photosynthetic zooxanthellae that lives within the polyp’s endodermis

          • Zooxanthellae = an algae

          • Endodermis = an inner layer of cells in the cortex of the root

        • The Zooxanthellae cannot survive in lower ocean temperatures

          • Below 18 degrees Celsius

        • Increases in ocean temperature cause Zooxanthellae to leave the coral tissue → this leads to coral bleaching

          • Above 35 degrees Celsius

        • Reef-building coral species therefore have a typical optimal growth range in temperate waters between 20 and 30 degrees Celsius

          • This correlates to the tropical and sub-tropical regions of the world (near the equator)

  • Species distribution

    • S: Use of a transect to correlate the distribution of plant or animal species with an abiotic variable

      • Quadrats and transects can measure the distribution of a plant or animal species in response to an incremental abiotic factor

        • Incremental = happening gradually

        • Quadrats = rectangular frames of known dimensions than can be used to establish population densities

        • Transects = a straight line along an abiotic gradient from which population data can be recorded to determine a pattern

        • Quadrats and transects are used together

      • Quadrats are placed at regular intervals along the transect line to generate population data

        • Basically, placing a quadrat down regularly and observing/counting the difference in distribution of X species based on the change of Y abiotic variable

        • The quadrats will show the changing distribution pattern of a species in response to a change in an abiotic variable

        • This data can be used to identify optimal conditions + zones of stress and zones of tolerance

      • Sampling with Quadrats and Transects

        • Quadrat counting method: count all variables mostly inside the quadrat (not mostly out)

        • Line transects: find a line, place quadrat every x meters

        • Transect data

          • Transects are used to assess species distribution in correlation with any abiotic factor that varies across a measurable distance

            • These factors could include elevation, elemental exposure, temperature, light levels, pH, humidity and more

        • A kite graph with the transect data can be used to represent changes in species distribution in a clear and effective fashion

          • The relative width of each kite represents the abundance of an organism at a particular point along a transect

          • Example:

  • Ecological niche

    • U: Each species plays a unique role within a community because of the unique combination of its spatial habitat and interactions with other species

      • Ecological niche = the functional position and role of an organism within its environment

        • Consists of all physical and biological conditions which determine the organism’s survival and reproductive prospects

        • Is comprised of various components, including:

          • The habitat in which an organism lives

          • The activity patterns of the organism

            • E.g. periods of time during which it is active

          • The resources it obtains from the environment

            • E.g. food sources, territorial boundaries, etc.

          • The interactions that occur with other species in the community

            • E.g. predator prey relationships, competition, etc.

    • U: Two species cannot survive indefinitely in the same habitat if their niches are identical

      • If two distinct species share an identical niche, there will be interspecific competition for available space and resources

        • This competition will result in the fitness of one being lowered by the presence of the other

        • The less well-adapted species will struggle to survive and reproduce

          • Eventually it will be eliminated from the niche

      • Interspecific competition within a shared niche will typically prompt one of two responses

        • Competitive exclusion → one species uses the resources more efficiently, driving the other species to local extinction

        • Resource partitioning → both species alter their use of the habitat to divide resources between them

          • Niche separation

      • Niche differentiation diagram:

    • S: Analysis of a data set that illustrates the distinction between fundamental and realized niche

      • Some species may not be able to occupy their entire niche due to the presence or absence of other species

        • Hence, a species may occupy a small subset of their niche than is theoretically possible

        • This creates fundamental vs. realized niche

      • Fundamental niche = the entire set of conditions under which an organism can survive and reproduce (where it could live)

        • The theoretical habitat

        • Example: in a rocky shore environment, the Chtalamus barnacle can potentially occupy the entire rocky shore, if in isolation

      • Realized niche = the set of conditions used by an organism after including interactions with other species (where it does live)

        • The actual habitat that is completely occupied by an organism in the presence of competing species

        • Example: the Chtalamus barnacle only occupies regions where the Semibalanus barnacle is absent

      • Diagrams:

        • Niches

        • Experimental data

        • Types of barnacles

  • Species interactions

    • U: Interactions between species in a community can be classified according to their effect

      • In nature, no species exist in total isolation → all organisms interact with both the abiotic environment and other organisms

        • The interactions between species can be classified according to their effect on the organisms involved

      • Herbivory

        • The act of eating only plant matter

          • That’s why primary consumers are called herbivores

          • Herbivores ay employ different feeding strategies

            • E.g. mucivores feed on plant sap while granivores feed on seeds, etc.

        • Herbivory can be either harmful or beneficial to plant species as a whole

          • Harmful: Certain types of beetle may feed voraciously on the leaves which causes crop failure

          • Beneficial: Fruit-eating animals spread the seeds from a fruit in their feces, which promotes overall seed dispersal

        • Examples of herbivores: Rabbit, cow

      • Predation

        • A biological interaction whereby one organism hunts and feeds on another organism

          • Predator and prey

        • The dependance of the predator on the prey as a food source intertwines their population levels

          • If the prey population drops, the predator numbers will fall as well as intraspecific competition has to increase

          • If the prey population rises, predator numbers will increase as a result of the ober-abundance of a food source

            • No intraspecific competition necessary = even the weakest survives

        • Diagram:

      • Symbiosis

        • Close and persistent long-term interaction between two species

          • Can be obligate (required for survival)

          • Or facultative (advantageous without being strictly necessary)

        • Symbiotic relationships can be beneficial to either one or both organisms in the partnership

          • Mutualism – both species benefit from the interaction

            • Example: how anemone protects clownfish and clownfish provides fecal matter for food

          • Commensalism – one species benefits, the other is unaffected

            • Example: barnacles are transported to plankton-rich waters by whales

          • Parasitism – one species benefits to the detriment of the other species

            • Example: ticks or fleas feed on the blood of their canine host

          • Diagram:

    • A: Local examples to illustrate the range of ways in which species can interact within a community

      • Mutualism

        • Ongoing interaction between two species whereby both species benefit from the interaction

        • Examples:

          • Honey bees gather nectar from flowers and distributes pollen between plants

            • This mediates the plant life cycle

          • Plover birds pick food morsels from between the jaws of crocodiles, cleaning their teeth in the process

          • Zooxanthellae photosynthesise within the protective environment of the polyp’s endodermis

            • This feeds the coral

      • Commensalism

        • Ongoing interaction between two species whereby one benefits and the other is unaffected

        • Examples:

          • Remora attaches to the underside of larger predatory fish and feed off the uneaten food scraps

          • Monarch butterflies can safely store poisonous chemical produced by milkweeds discourage predators from eating it

          • Decorator crabs remove small fragments of tissue from sea sponges and uses them as a source of camouflage

      • Parasitism

        • Ongoing interaction between two species whereby one species benefits at the other’s expense

        • Examples:

          • Ticks infest the skin and fur of host animals, feeding off the host and potentially causing disease

          • Leeches attach to the skin and drinks the blood of the host animal until fully engorged

          • Tongue-eating louses eat the tongue of a fish and may still later ingested food

    • A: The symbiotic relationship between Zooxanthellae and reef-building coral reef species

      • Reef-building coral will form a mutualistic symbiotic relationship with the photosynthetic unicellular algae Zooxanthellae

        • Coral are colonial organisms made up of individual polyps that are connected by a layer of living tissue

        • The algae lives within the cells of the coral’s endodermis (the innermost lining of the animal)

      • The coral provides the algae with a protective environment and source of inorganic compounds

        • Coral polyps secrete calcium carbonate to build a stony skeleton which encases the polyps and the Zooxanthellae

        • Coral polyps also recycle the waste products of the algae and supplies the Zooxanthellae with CO2

      • The Zooxanthellae in turn provides the coral polyps with a necessary source of nutrition

        • The alage supplies the coral with oxygen, glucose and other organic molecules that it produces via photosynthesis

        • The algae also helps the coral remove necessary waste products

      • Diagram of their mutualistic relationship:

      • Coral bleaching

        • It is the Zooxanthellae that gives the coral its vibrant pigmentation

        • When there is a large scale loss of Zooxanthellae from the coral due to environmental stress, bleaching of the coral occurs

          • This causes the coral to starve and die unless the Zooxanthellae are restored

        • Conditions which can cause coral bleaching include:

          • Changes in light availability

            • E.g. sedimentation may increase the opacity of the oceanic waters

          • Temperature increases

            • Water temperatures in excess of 30 degrees Celsius can irrevocably stress the Zooxanthellae

          • Ocean acidification

            • The buildup of CO2 concentrations in the ocean can lower pH and stress the Zooxanthellae

  • Keystone species

    • U: Community structure can be strongly affected by keystone species

      • Keystone species = a species that has a disproportionately large impact on the environment relative to its abundance

        • Name comes from the keystone in an arch → it fundamentally supports the whole structure and prevents it from collapsing

      • Keystone species may influence communities in a number of ways:

        • Predators can exert pressure on lower trophic levels to prevent them from monopolizing certain resources

        • Mutualists can support the life cycle of a variety of species within a community

          • E.g. pollinators and seed dispersal

        • Engineers can refashion the environment in a manner that promotes the survival of other species

          • Engineer = any species that creates, significantly modifies, maintains and/or destroys habitats

      • Numerous examples of keystone species within different communities, such as:

        • Sea stars (predators) prey on urchins and mussels, which prevents mussel overpopulation and coral reef destructions by urchins

        • Honey bees (mutualist) pollinate a wide variety of plant species, ensuring the continuation of the plant life cycle

        • Beavers (engineer) build dams that transform the environment in a manner that allows certain other species to survive

      • Keystone species are not the dominant species (most numerous) within a community nor do they have to be apex predators

      • Diagram of consequences of removal of keystone species:


C2 Communities and ecosystems

  • Trophic levels

    • U: Most species occupy different trophic levels in multiple food chains

      • Trophic level = the position an organism occupies within a feeding sequence

        • Producers occupy the first trophic level in a feeding sequence

        • Primary consumers feed on producers → second trophic level

        • Secondary consumers feed on primary → third trophic level

        • Further consumers occupy the subsequent levels

        • Diagram:

      • Food chain = shows linear feeding relationships between species in a community

        • Arrows represent the transfer of energy and matter

          • This transfer happens as one organism is eaten by the other

          • The arrows point in the direction of energy flow

        • Goes from the producer to the consumers

        • Most species’ feeding requirements means they will be in multiple food chains where they may occupy a different trophic level

          • As few species only feed on one other species

        • E.g.:

    • U: A food web shows all the possible food chains in a community

      • Links food chains together into more complex and interrelated feeding relationships

      • More representative of actual feeding pathways within an ecosystem because

        • Organisms can have more than one food source

        • Organisms can have more than one predator

        • Hence, organisms can also occupy more than one trophic level

      • Tip! When constructing food webs, position organisms at their highest trophic level (from left to right) as it keeps all arrows pointing in the same directions

      • Example:

    • EC: Ecological Pyramids

      • Ecological pyramids show the relative amounts of a specific component at the different trophic levels of an ecosystem

        • The three main types measure species numbers, biomasses and energy

      • Pyramid of Numbers

        • Shows the relative number of organisms at each stage of a food chain

        • Are usually shaped like pyramids → higher trophic levels cannot be sustained if there are more predators than prey

          • However, the shape may be distorted if a food source is disproportionately large in size/biomass compared to the feeder

            • E.g. a large number of caterpillars may feed on a single oak tree

      • Pyramid of Biomass

        • Shows the total mass of organisms at each stage of a food chain

        • Almost always upright in shape as biomass diminished along food chains as CO2 and waste is released

          • An exception to this rule is found in marine ecosystems where zooplankton have a larger total biomass than phytoplankton

            • Because phytoplankton replace their biomass at such a rapid rate and can therefore support a larger biomass of zooplankton

      • Pyramid of Energy

        • Shows the amount of energy trapped per area in a given time period at each stage in a food chain

        • Always upright in shape as energy is lost along food chains

          • Either used in respiration or lost as heat

        • Each level in the pyramid is roughly one tenth the size of the preceding level as energy transformations are ca. 10% efficient

      • Examples of pyramids:

  • Energy conversions

    • U: The percentage of ingested energy converted to biomass is dependent upon the respiration rate

      • Ecological production/productivity refers to the rate at which biomass is generated in an ecosystem

        • Usually expressed in units of mass per area per time

          • km m^-2 day^-1

      • Biomass = total dry weight of organic matter in organisms or ecosystems

        • The entirety of all biologically produced organic matter

          • E.g. proteins, carbohydrates, etc.

      • The percentage of energy that is converted into biomass is dependent upon a number of factors that lose energy

        • Energy is lost as inedible materials (bones, teeth, hair)

        • Energy is lost via excretion of undigested or unabsorbed materials

        • Energy is lost as heat from cellular respiration

          • The higher respiration rate results in more heat lost

      • Primary production = the production of chemical energy in organic compounds by producers

        • The main source of energy for primary production is sunlight

          • Sometimes chemosynthesis by lithotrophs

        • Can be categorized as one of two types:

          • Gross primary production (GPP) → the amount of chemical energy as biomass that a producer creates in a given length of time

          • Net primary production (NPP) → the amount of chemical energy that is not consumed by respiration

            • GPP - respiration

      • Secondary production ? the generation of biomass by heterotrophic organisms (consumers)

        • Biomass generation is driven by the transfer of organic compounds between trophic levels via feeding

        • Secondary production may also be categorized according to gross and net amounts of biomass

          • Gross = total

          • Net = usable

      • Diagram explaining how ingested energy converted to biomass depends on respiration:

  • Feed Conversion Ratio

    • A: Conversion ratio in sustainable food production practices

      • Feed Conversion Ratio (FCR) = mass of feed/mass of desired output

        • FCR measure the efficiency of an animal in converting food into a desired output

        • The lower the FCR the more efficient the method of food production

      • Low FCR is achieved by minimizing the potential losses of energy for the animal stock

        • Restricting animal movement will reduce energy lost to cell respiration

        • Optimizing feeding practices so that food is ingested and digested more effectively

        • Slaughtering animals at a young age

          • As older animals tend to grow more slowly and have a higher FCR

      • Theoretically the most efficient FCR is best as it lowers cost, however there are a number of potential ethical issues involved

        • Is it ethical to restrict animals and kill them early?

      • Diagram displaying the FCRs of different species:

    • EC: Sustainable food practices

      • Most of the protein found in Western human diets comes from farmed animal meet

        • E.g. cattle and poultry

        • These animals must be fed and housed → this creates an ecological footprint, the size of which depends on the FCR

      • Sustainable food practices provide food for the consumer while minimizing the environmental impact of the food source

        • The benefits of lowering food production costs must be balanced against ethical considerations regarding animal treatment

      • One sustainable food practice is the use of crickets as a protein source (as opposed to cattle)

        • Crickets have a substantially lower FCR → minimizes feed costs

        • They require significantly less water → minimizes water usage

        • They can reproduce much faster and require less space

        • Produce far less methane

        • For these reasons, certain health professionals are advocating the introduction of crickets into normal diets

          • The advantages of crickets as a food source is still debated though, so implementation is likely to be slow

  • Types of Ecosystems

    • U: In closed ecosystems energy but not matter is exchanged the with the surroundings

      • Ecosystems can either be open or closed, depending on whether matter moves into and out of the system

        • A closed ecosystems exchanges energy with its surrounding, but not matters

          • It is self-contained

          • E.g. a mesocosm

        • An open ecosystem changes both energy and matter with surrounding environments

          • A natural ecosystem such as a forest

    • U: The type of stable ecosystem that will emerge in an area is predictable based on climate

      • Ecosystem = the interaction of living and non-living things within an area

        • A community and its abiotic environment

      • Particular types of stable ecosystems will emerge in a given geographical area according to its climate conditions

      • Biome = a geographical area that has a particular climate and sustains a specific community of plants and animals

        • Basically a specific type of natural ecosystem, created from its climate and environment

        • The main factors affecting the distribution of biomes is temperature and rainfall

        • These factors vary according to latitude, longitude, altitude, proximity to ocean, etc.

      • Many different biomes are found across continents, including:

        • Tropical rainforests

          • Hot and humid environments

          • Near the equator

          • Dense vegetation

          • High biodiversity

        • Taiga

          • Coniferous forests

          • Near the poles

          • Have cold temperatures

          • Little precipitation

            • Moisture trapped as snow or ice

        • Deserts

          • Dry and arid environments

          • Display extreme temperature conditions (hot and cold)

        • Picture examples:

    • EC: Types of Biomes

      • Biomes are geographical area that have a particular climate and sustain a specific community of flora and fauna

        • The total area where all living things are found is called the biosphere

          • The totality of biomes

        • Certain biomes are named and recognized according to their specific characteristics

      • Tropical rainforest

        • Hot climate → 25-30 degrees Celsius

        • Very high levels of precipitation → > 250 cm per year

        • Large diversity in species

        • Vegetation includes epiphytes, tall trees and undergrowth

          • Epiphytes: organisms that grow on the surface of plants

      • Temperate forest

        • Also called deciduous forests

        • Moderate temperatures and clear seasonal changes

        • Growing period of around 200 days during 4-6 frost free months

      • Boreal forest (Taiga)

        • Coniferous forests

        • Cold and icy → 0-15 degrees Celsius

        • Only small amount of precipitation

        • Coniferous trees are densely packed and there is little variation in species

      • Tropical Grassland (Savannah)

        • Warm → 20-30 degrees Celsius

        • Medium amounts of moisture, although seasonal droughts are common

        • Trees are intermittent and widely spaced

        • Occasional fires prevents thicker vegetation from encroaching

      • Temperate Grassland

        • Moderate temperatures and amounts of rainfall

        • Trees and shrubs are largely absent

        • Grass is the dominant vegetation

      • Mediterranean (Chaparral)

        • Moderate temperatures → 15-25 degrees Celsius

        • Rainy winters and dry summers

        • Vegetation consists of dry woody shrubs that are quick to grow

      • Desert

        • Extreme temperature conditions → > 30 degrees Celsius during the day, < 0 degrees at night)

        • Very low precipitation

          • < 30 cm per year

        • Dominant plant species are xerophytes which are adapted for water conservation

      • Tundra

        • Freezing temperatures → < 0 degrees Celsius

        • Very little precipitation

        • Vegetation is low growing

          • E.g. mosses

        • Perennial plants may grow during the summer

      • Mountain ranges

        • Found all over the world at high attitudes → > 10 000 feet

        • Temperatures are typically low due to the altitude

        • Weather conditions may change rapidly

      • Distribution of biomes in the biosphere:

  • Ecosystem Analysis

    • S: Analysis of a climograph showing the relationship between temperature, rainfall and the type of ecosystem.

      • Climograph = graphical representation of basic climate parameters at a given geographical location

        • Shows a combination of monthly average temperature and precipitation at a certain location

          • Precipitation = rainfall

        • Provides a quick overview of the climate of a region and can be used to identify seasonal patterns and changes

        • Examples:

      • Whittaker’s climograph

        • In 1975, ecologist Robert Whittaker developed a holistic climograph that allowed for the classifying of 9 distinct biomes

        • Biomes distinguished according to their average yearly temperatures and precipitation

          • Deserts typically have high average temperatures but low precipitation

            • Hot and dry

          • Rainforests typically have high average temperatures and high precipitation

            • Hot and wet

          • Taigas typically have low average temperatures and reasonably low precipitation

            • Cold and icy

    • S: Comparison of pyramids of energy from different ecosystems

      • Pyramids of energy (PoE) = diagram of the flow of energy between trophic levels

        • Measures in units of energy per area per time

        • Standard units are kJ m^-2 and y^-1

        • Triangular and never inverted as 90% of energy is lost between trophic levels

      • PoEs differ between ecosystems due to the effect of climate on primary productivity

        • Warmer temperatures speed up enzyme reactions required for photosynthesis

        • High precipitation also increases photosynthesis as the photolysis of water is essential for non-cyclic photophosphorylation

        • Higher (optimal) rate of photosynthesis = more energy transferred

      • Tropical rainforests have a high NPP whereas deserts have a low NPP

        • Higher NPP = more energy supplies to consumers hence more trophic levels can exist

        • Tropical rainforests PoE has a wider base and more levels than a desert

      • Types of species within a particular biome may also affect how efficiently energy is transferred between trophic levels

        • Example: Homeotherms will use more energy maintaining a stable body temperature compared to poikilotherms

      • Diagram:

    • S: Construction of Gersmehl diagrams to show the interrelationships between nutrient stores and flows between taiga, desert and tropical rainforest

      • Gersmehl diagrams show the differences in nutrient flow and storage between different types of ecosystems

      • Nutrients are stored in one of three nutrient sinks

        • Biomass → the total mass of living organisms in a given area

          • Mainly plant tissue

        • Litter → any organic matter in and on the soil

          • Includes humus and leaf litter (defoliation)

        • Soil → the top layer of the earth that is composed of disintegrated rock particles

      • Nutrients can be transferred between nutrient sinks and may also be cycled via environmental inputs and outputs

        • Transfer from biomass to litter → fallout

        • Transfer from litter to soil → decay

        • Transfer from soil to biomass → uptake

        • Litter can additional gain nutrients via precipitation and lose nutrients in surface runoff

        • Soil can gain nutrients from the erosion of rocks via weathering and lose via leaching

      • Standard: Gersmehl diagram

      • Comparing Gersmehl diagrams

        • The inter-relationships between nutrient stores and flows will be affected by climatic factors

          • Such as temperature and rainfall

          • Therefore Gersmehl diagrams will differ between biomes

        • Tropical rainforest

          • Most nutrients are stored as biomass

            • Litter is rapidly decomposed and vast roots quickly draw nutrients from soil

          • There is a fast rate of transfer between stores

            • Hot and wet conditions promote precipitation, runoff, weathering and leaching

        • Taiga

          • Most nutrients are stored as litter

            • Low temperatures slow decomposition which delays nutrient transfer to soil and biomass

          • There is little nutrient gain from precipitation or weathering due to low levels of precipitation

            • Cold temperatures produce snow instead

          • There is little nutrient loss via leaching due to low precipitation

            • But surface runoffs may be high at certain times if the snow melts

        • Desert

          • Most nutrients are stored in the soil

            • Few plants exist to store nutrients as biomass nor to produce litter

          • There is little nutrient gain from precipitation and little nutrient loss via runoff → both due to the dry climate

          • The amount of weathering and leaching is insignificant

  • Primary Succession

    • S: Analysis of data showing primary succession

      • Ecological succession = the process by which a sequence of increasingly complex communities develop over time

        • Climax community = when succession has ended and the community has all of its characteristics

      • Primary succession = when communities develop on entirely new land without any established soil

        • May occur at river deltas, glaciers, sand dunes or on exposed rock

        • First organisms to colonize the region are called pioneer species

          • Typically consist of lichen or moss

            • When they die, they decompose which creates the first organic soil capable of sustaining plant growth

      • As plant species colonize a new area, the litter produced by their growth and decomposing remains will cause changes

        • Soil depth will increase because plants add humus to the soil

        • Soil pH will become altered by the additions

        • Soil mineral content will increase and rocks will begin to be broken down by the action of roots

        • The soil will become aerated and water retention increases

          • Drainage is reduced

        • These changes will allow for growth of larger plants, which will reduce erosion through the binding action of their roots

          • Not all plant species will thrive in unison → larger plants will eventually outcompete smaller shade intolerant plants

      • Overview of primary succession:

      • Primary succession data

        • A primary succession sequence can be identified according to the distribution of plants

          • This applies for any location where a new community may emerge from uninhabitable land

          • The regions closer to the site of development will be in the earlier chronological stages of succession

            • E.g. Glacial retreat exposes bare rock for colonization, so regions further from the glacier have had more time to develop life

        • Diagram example:

  • Ecological Disturbance

    • U: Disturbance influxes the structure and rate of change within ecosystems

      • Environmental disturbances may cause fluctuations to the structure and rate of change within ecosystems

        • Such disturbances may be natural or artificial (human-induced)

          • E.g. bushfire, earthquake

      • Natural disturbance

        • A natural environmental disturbance may give rise to secondary succession

          • Secondary succession = one ecosystem is replaced by another

          • Occurs when succession starts on existing soil following the upheaval (change) of a pre-existing ecosystem

            • This upheaval results in the removal of existing biota and allows a new ecosystem to develop on the site of the old

          • Because the soil is already developed during secondary succession, dominance is usually achieved by the fastest growing plants

        • The progression of secondary succession can be summarized as follows:

          • An environmental disturbance destroys the pre-existing climax community

          • Grasses and herbaceous plants are the first to grow back as the soil is already present

            • Soil = no pioneer species required

          • Fast growing trees will develop to their fullest while shade tolerant trees will develop in the understory

            • Eventually the fast-growing trees may be overtaken by larger, slower-growing trees as the ecosystem reverts to its prior state

        • Diagram:

    • A: Consideration of one example of how humans interfere in nutrient cycling

      • Deforestation is the permanent destruction of a forest via the removal or clearance of trees

        • Human-induced degradation of forests

        • Driven primarily by a need for timber and cleared land for agricultural purposes

      • Deforestation disturbs the normal nutrient cycling within the region in a number of ways

        • Less trees means there is less evapotranspiration  meaning there is less moisture/precipitation in the air

        • Fewer trees means less litter

          • Due to less defoliation (leaves falling off a plant)

          • This reduces the production of humus, so there will be less nutrients in the soil

        • Rapid loss of nutrients from leaching

        • Less chemical weathering of rock, due to less water in soil

        • The soil will become more acidic and release iron and aluminum to form an infertile ferralitic soil

          • Ferralitic soil = nutrient poor

        • The soil layer will become increasingly thin and eroded

        • The infertile soil will prevent vegetative growth, reducing biodiversity and nutrient cycling

        • Logging operations may also alter the distribution of plant species by removing the canopy and increasing light exposure

          • Canopy = the upper layer of habitat zone, usually formed by mature tree crowns

        • Removal of the canopy also results in an increases loss of nutrients from the soil via runoff

      • The effect of deforestation on nutrient cycling:

    • S: Investigation into the effect of an environmental disturbance on an ecosystem

      • Environmental disturbances are caused by natural or artificial disruptions to a normal ecosystem

        • Examples:

          • Fire breaks in bush lands or regions damaged by bushfires

          • Outer boundaries of population settlements or regions bordering roads

          • Dams and artificial rivers and creeks

            • E.g. irrigation sites

      • The effect of an environmental disturbance on an ecosystem can be measured in a number of ways

        • Population density

          • Using the Lincoln index via the capture-mark-recapture technique

        • Species diversity and richness

          • Using the Simpson’s reciprocal index

        • The presence and distribution of indicator species

          • To measure levels of pollution

        • Canopy coverage and relative light intensity

          • Could measure with a lux meter

        • Biomass

          • Via the average width of tree stems at a specified height

        • Edaphic factors such as soil erosion (via depth), water retention (via drainage), pH and nutrient content

      • Measurements taken from a disturbed area need to be compared against measurements taken from an undisturbed control area

        • This enables the investigator to statistically calculate both the effect and magnitude of the environmental disturbance

C3 Impacts of humans on ecosystems

  • Invasive species

    • U: Introduced alien species can escape into local ecosystems and become invasive.

      • The species within any given ecosystem can be broadly described as either endemic or alien

        • Endemic = native species to a defined geographic region

          • E.g. koalas in Australia

        • Alien = species that have been transferred from their natural habitat into a new environment

        • If an introduced alien species has detrimental effects on the pre-existing food chains, it is classified as invasive

          • Invasive species typically threaten the biodiversity of the ecosystem they are introduced into by displacing native species

      • Example: invasive species in Australia → cane toad, European rabbit, red fox, camel, feral cat

    • U: Competitive exclusion and the absence of predators can lead to reduction in the number of endemic species when alien species become invasive

      • Competitive exclusion principle = two species cannot occupy identical niches within a community

        • One species will have a competitive advantage and survive at the expense of the other

        • Evolution via natural selection results in adaptations that function to minimize direct competition between species in nature

      • Invasive species possess several advantages that allow them to displace endemic species via competitive exclusion:

        • Typically possess a large fundamental niche

          • = can occupy a wider area than endemic species

        • Commonly lack a predator capable of limiting their survival

          • As they are introduced into a foreign environment

        • May possess certain features that are suited to the new environment

          • E.g. more aggression, better foraging, etc.

      • Diagram:

    • A: Study of the introduction of cane toads in Australia and another local example of an introduced alien species

      • Case study one: Cane toads (Bufo marinus)

        • Cane toad was introduced in Australia in 1935 by humans to control the spread of the sugar cane beetle that was damaging crops

          • Failed to limit the beetle population → beetles live at the tops of sugar canes where the toads could not reach

        • The cane toad has reproduced rapidly and is now classified as an invasive species that endangers native wildlife

          • It has a wide diet → depleted the prey population for native insectivores

          • Secretes a toxic chemical from its skin that has poisoned native predators

          • Carries diseases that have been transmitted to native species of frogs and fish

        • The cane toad has spread across the northern fringes of Australia

          • Now numbers in the millions

        • Proven difficult to eradicate as there are no natural predators in Australia to reduce its population size

      • Case study 2: Wild rabbits

        • European rabbits were introduced in Australia with the arrival of the First Fleet

          • Ostensibly for recreational hunting

        • Australian conditions promoted a population explosion

          • Mild winters allowed rabbits to breed the entire year

          • Fastest spread ever recorded for any mammal anywhere in the world

        • Rabbits have had a devastating effect on native Australian ecology

          • Killed native plant species by ringbarking

            • Ringbarking = removing a strip of bark that prevents future growth beyond that point

          • Loss of native plants has left the topsoil exposed and vulnerable to extensive erosion from wind and rain

        • Various strategies have been employed to control rabbit populations with varying effect

          • ‘Rabbit-proof’ fences were built to try and contain rabbit populations

          • Methods of hunting and poisoning have been utilized

            • Also efforts to destroy rabbit warrens (burrows)

          • Rabbit-borne diseases have been released with moderate success

            • E.g. myxomatosis virus

  • Biological control

    • S: Evaluation of eradication programmes and biological control as measures to reduce impact of alien species.

      • Controlling the population of invasive species once they have become established is difficult and typically very expensive

        • Three main methods: physical, chemical, biological

      • Physical control

        • Involves removal or restriction by manual or mechanical measures

        • May include:

          • Installation of barriers and fences

          • Removal of habitat by excavation or trimming

          • Population numbers may be reduced by hunting, trapping and culling

            • Although this is labor intensive

        • Physical methods to contain invasive species are not usually species specific and can also impede endemic wildlife

      • Chemical control

        • Involves the use of chemical agents to limit population numbers and spread

          • Chemical agents = poisons and toxins

        • Chemical agents may include:

          • Herbicides for plants

          • Pesticides for insects

          • Other compounds such as rat poison

        • Chemical agents may have moderate specificity, but can also detrimentally affect local wildlife + are costly

        • The effect may become more pronounced in high trophic levels due to biomagnification

      • Biological control

        • Involves using living organisms or viruses to control an invasive species

        • Biological control may eat the invasive species or cause it to become diseased

        • Biological agents mut be carefully assessed before release to ensure they do not become invasive themselves

        • Examples:

          • The Vedalia beetle feeds on citrus plant invertebrates

          • The Myxoma virus that affects rabbits

        • Biological control agents must be monitored for unintended side effects

          • E.g. development of immunity in invasive species

        • Biocontrol case study: The Vedalia Beetle (ladybug)

          • The cottony cushion scale is an invertebrate pest from Australia that was accidentally released in California

            • Spread and fed on citrus plants such as orange trees

            • Devastated the Californian citrus industry

          • The Vedalia beetle is a predatory insect from Australia that was introduced into California as a means of biological control

            • It worked to limit the numbers of the cottony cushion scale

            • Minimized the economic impact on the citrus industry

          • Diagram:

  • Biomagnification

    • U: Pollutants become concentrated in the tissues of organisms at higher trophic levels by biomagnification

      • Biomagnification = the process in which chemical substances become more concentrated at each trophic level

        • Bioaccumulation = the build up of a chemical substance in the tissues of a single organism

      • Biomagnification occurs because organisms at higher trophic levels must consume more biomasse to meet feeding requirements

        • Energy transformations are only ca. 10% efficient, so higher order consumers must eat more to meet energy demands

        • This means higher order consumers will experience increased contamination from a chemical substance

      • Diagram of effect:

    • S: Analysis of data illustrating the causes and consequences of biomagnification

      • Dichloro Diphenyl Trichloroethane (DDT) is an example of a chemical substance which is biomagnified

        • DDT = a chemical pesticide that is sprayed on crops to eliminate mosquito larvae and washed into waterways at low concentrations

          • Taken up by algae and passed on to primary consumers

        • DDT is fat soluble and selectively retained within the tissues of an organism instead of being excreted

      • At each subsequent trophic level DDT is passed on through the algae, the concentration of DDT stored in the body increases

        • Increased trophic level = increased food uptake = increased DDT

        • Very high levels of DDT were discovered in birds that preyed on fish

      • Birds exposed to high levels of DDT through feeding on fish that fed on algae were found to produced thinner shells as a consequence

        • This decreased survival rates of fledglings

      • Diagrams we may encounter for such data analysis

    • A: Discussion of the trade-off between control of the malarial parasite and DDT pollution

      • DDT was widely employed as a chemical pesticide in the 1940s and 1950s to control insect-borne diseases like malaria

        • Agricultural insecticide

        • Led to complete eradication of malaria in many high socioeconomic world regions

        • Less effective in low socioeconomic tropical regions where eradication programs could not be sustained

      • Eradication programs were curtailed by the World Health Organization (WHO) in 1969

        • For safety and environmental concerns

        • DDT can bioaccumulate to toxic levels and poses a significant threat to high trophic levels due to biomagnification

        • Current interventions for malaria now focus on non-spraying strategies

          • E.g. the use of bednets soaked in insecticides

      • Malaria vs. DDT

        • Arguments for DDT spraying:

          • DDT spraying is an affordable and effective means of killing mosquitos that carry disease

            • E.g. malaria, dengue fever, etc.

          • Where the use of DDT has been discontinued, the incidence of malaria and associated deaths have increased

          • Health costs associated with the treatment of malaria are reduced when DDT spraying is implemented

          • Alternative strategies are not as cost-effective or successful

        • Arguments against DDT spraying

          • It is associated with adverse health effects in humans

            • E.g. cancer, birth defects, reduced fertility, etc.

          • Persists in the environment for long periods of time

            • More than 15 years

          • DDT is biomagnified in higher order consumers which has adverse consequences on ecosystems

  • Plastic pollution

    • U: Macroplastic and microplastic debris has accumulated in marine environments

      • Plastic = a type of synthetic polymer

        • Found in certain types of clothes, bottles, bags, food wrappings, containers, etc.

        • Most plastics are not biodegradable → persist in the environment for many centuries

      • Macroplastic = large visible plastic debris

        • 1 mm

        • Macroplastic debris can be degraded and broken down into microplastic debris by UV radiation and the action of waves

      • Microplastic = smaller plastic debris

        • < 1 mm

      • Ocean currents will concentrate plastic debris in large oceanic convergence zones called gyres

      • Plastic debris in the ocean will leach chemical into the water + absorb toxic contaminants called persistent organic pollutants

        • Microplastics will absorb more persistent organic pollutants (POPs) due to their smaller size

          • More surface area available

      • Both macro- and microplastic debris is ingested by marine animals, which mistake the debris for food

        • This leads to the bioaccumulation and biomagnification of POPs within marine animals

        • It may also damage the stomach of animals or cause them to stop feeding

          • By taking up space in the digestive tract

      • Diagram:

    • A: Case study of the impact of plastic debris on Laysan albatrosses and one other named example

      • Case Study 1: Laysan Albatross

        • Nests on islands found in the North Pacific gyre where large amounts of plastic debris is found

        • Feeds by skimming the ocean surface with their beak → this causes them to ingest large quantities of plastic too

        • Adults can regurgitate (throw up) the plastic they have swallowed, but chicks are unable to

          • Instead, it fills up their stomachs

          • The mortality rate in albatross chicks is very high

            • Estimated 40% die before fledgling

      • Case Study 2: Sea turtles

        • Will commonly mistake plastic bags for jellyfish

          • Jellyfish = one of their primary food sources

        • Ingestion of the plastic can be fatal

          • It can become lodged in the esophagus and cause future feeding problems

        • Plastic debris can also become wrapped around the turtle → restricting movement and developmental growth

        • It is estimated plastic pollution harms ca. 100 000 sea turtles and other marine animals each year

  • EC: Cholorfluorocarbons

    • Chlorofluorocarbons (CFCs) = chemicals widely used as refrigerants, propellants and solvents in the past

      • Contribute to ozone depletion in the upper atmosphere

        • Therefore their manufacture and use is being phased out

    • Ozone layer = stratospheric region composed of ozone (triatomic oxygen)

    • CFCs are broken down by UV radiation to release chloride ions that react with ozone

      • The ions breaks the ozone down into oxygen

      • One chloride ions can destroy 100 000 ozone molecules

      • Diagram of the process:

    • Ozone in the stratosphere absorbs UV radiation, however there is a limit to how much radiation ozone can absorb

      • Regions where the ozone layer is thinner will experience higher terrestrial UV levels

        • Thinning caused by for example CFCs

      • UV light penetrates tissues and damages DNA → this causes gene mutations and cancer

      • Can also damage the ability of plants to carry out photosynthesis + kill phytoplankton → this reduces productivity

  • EC: Earth spheres

    • The Earth is made up of four distinct and connected spheres

      • Lithosphere, hydrosphere, atmosphere, biosphere

    • Lithosphere = all the rocks, minerals and molten magma found on or in the Earth

      • Also called geosphere

    • Hydrosphere = all the water on Earth

      • Liquid water such as oceans and rivers

      • Precipitation and vapor

    • Atmosphere = the layer of gasses surrounding the Earth

      • Divided into further sections

      • E.g. stratosphere

    • Biosphere = all the living organisms on the planet

      • Including plants, animals, bacteria, etc.

    • As the four spheres are interconnected, human impact on one sphere will potentially affect other spheres

      • Examples:

        • The release of plastic pollution into the hydrosphere (oceans) will impact the biosphere (marine life)

        • The production and release of CFCs into the atmosphere will impact how much UV radiation the biosphere is subjected to

C4 Conservation of biodiversity

  • Indicator species

    • U: An indicator species is an organism used to assess a specific environmental condition

      • Indicator species are sensitive to specific environmental conditions → limited range of tolerance

        • Reduction or growth of their population therefore indicates changes in the environment

          • Useful means of monitoring change

      • Examples of indicator species and their environmental conditions

        • Lichen and mosses are susceptible to air-borne pollutants dissolved in water

          • E.g. sulfur dioxide

        • Tubifex worms are sensitive to concentrations of heavy metals

        • Mayfly larva and certain aquatic invertebrates are sensitive to dissolved oxygen levels in water

      • Diagram:

    • U: Relative numbers of an indicator species can be used to calculate the value of a biotic index

      • Biotic indices compare the relative frequency of indicator species

          • Provide an overall assessment of environmental health

      • Calculating a biotic index involves multiplying the population size of each indicator species by its pollution tolerance rating

      • The following conclusions can be drawn from a biotic index:

        • A high biotic index = the presence of many pollution-sensitive organisms

          • Denotes an unpolluted environment

        • A low biotic index = the absence of pollution-sensitive organisms + abundance of pollution-tolerant organisms

          • Indicates a polluted environment

        • A change in the biotic index over time marks a change in the environmental conditions within a given ecosystem

      • Pollution tolerance rating

      • Environmental pollution levels

  • Biodiversity

    • U: Richness and evenness are components of biodiversity

      • Biodiversity = the variety and variability of all living organisms within a given ecological area

        • Can be used to refer to the number of species, their genetic diversity or habitat variety

          • Habitat variety = ecological variations

      • 2 main components contribute to biodiversity

        • Species richness → the number of different species present in an area

          • More species = greater richness

        • Species evenness → the relative abundance of the different species in an area

          • Similar abundance = more evenness

        • Richness vs. evenness

    • S: Analysis of the biodiversity of two local communities using Simpson’s reciprocal index of diversity

      • Simpson’s reciprocal index can be used to measure the relative biodiversity of a given community

        • It takes into account both the number of species present (richness) and the number of individuals per species (evenness)

        • A higher index value is indicative of a greater degree of biodiversity within the community

      • Simpson’s reciprocal index can be used to compare communities to identify intrinsic values

        • Intrinsic value = something we value for itself, not its consequences

        • A high index value = a stable site with many different niches and low competition

          • High richness and evenness

        • A low index value = a site with few potential niches where only a few species dominate

          • Low richness and evenness

        • The index value may change in response to ecological disturbance

          • E.g. human intervention, natural disasters, etc.

  • Biogeographic factors

    • U: Biogeographic factors affect species diversity

      • Biodiversity within a given ecosystem will be affected by a number of biogeographic factors

        • Larger habitats tend to promote biodiversity better than smaller ones

          • More available niches = less competition

        • Ecology at the edges of ecosystem is different from central areas

          • E.g. more sunlight, more wind, etc.

          • This is known as the edge effect, whereby species distribution is influenced by divergent environmental conditions

          • Edges tend to have greater biodiversity as different habitats with different abiotic factors exist in close physical proximity

          • However, edges tend to have more competition than central regions → may restrict survival prospects of certain species

        • Habitat corridors between parts of a fragmented habitat can connect disparate regions to improve genetic diversity

      • These principles of biogeographic factors are routinely applied when setting aside land as a nature reserve to improve the conservation of biodiversity

    • A: Analysis of the impact of biogeographic factors on diversity limited to island size and edge effect

      • Island size

        • The biodiversity of an island is typically proportionate to island size

          • Larger island = greater biodiversity

            • Larger islands support a greater range of habitats

              • Hence more available niches for species to occupy → more survival + diversity

            • Larger islands can sustain higher population numbers for each species

              • Increases species evenness

            • Larger islands have greater productivity at each trophic level

              • Leads to longer + more stable food chains

          • Smaller island = less biodiversity

          • Diagram depicting the relationship between island size and biodiversity

      • Edge effect

        • The diversity of species within a given environment changes with proximity to the ecosystem’s boundaries

        • Biodiversity may be greater at the border between two ecosystems as different abiotic factors favor certain species

        • However, certain species may not be able thrive under these conditions

          • Instead must occupy more central regions

        • The effect of edges on biodiversity will be greatly influenced by the particular conditions caused by the ecological disturbance

        • Diagram:

  • Conservation

    • U: In situ conservation may require active management of nature reserves or national parks

      • Conservation = the careful preservation, protection and maintenance of natural resources

        • Such as trees, water, wildlife, etc.

        • Can be either in situ or ex situ

          • On site or off site

      • In situ conservation is the preservation of plant and animal species within their natural habitat

        • Typically involves the designation of protected areas of land as either nature reserves or national parks

      • These areas may require active management to ensure that an appropriate and sustainable ecological balance is maintained

        • Ecological monitoring of a species may be required to ensure viable population levels are maintained

        • Interventions may be required to prevent habitat degradation or competition from invasive species

        • Legislation may be necessary to ensure adequate funding for policing and education

      • In situ conservation offers several advantages when protecting endangered species from extinction

        • It allows species to live in the environment to which they are adapted

          • + to occupy their natural position in the food chain

        • It maintains the animal’s normal behavior

          • Offspring usually acquire skills from parents and peers around them

        • Retaining the natural habitat prevents its eventual loss + ensures it remains available for other endangered species

          • Such areas provide a place to return animals from breeding programs as they provide realistic conditions for reintegration

        • Reserves in different areas can share information and provide a place for scientific study and developing public awareness

    • U: Ex situ conservation is the preservation of species outside their natural habitats

      • Ex situ conservation involves the preservation of plant and animal species outside their natural habitats

        • May typically be required for critically endangered species when urgent intervention is necessary

      • There are several advantages associated with ex situ conservation

        • It allows for greater control of essential conditions

          • E.g. climate control, dietary intake, veterinary care, etc.

        • It can improve the chances of successful breeding by utilizing artificial methods

          • E.g. embryo transfer, IVF, etc.

      • Ex situ conservation is also associated with several disadvantages

        • They do not prevent the potential destruction of their natural habitats

        • Species raised in captivity are less likely to be successfully reintroduced into the wild

          • Loss of autonomous survival

        • Increases inbreeding by restricting the gene pool → restricts the evolution of the species

      • Some examples of ex situ conservation measures employed around the world:

        • Captive breeding → animals are raised and bred in containment to ensure survival prospects

          • E.g. zoos

        • Botanical gardens → areas devoted to the collection, cultivation and display of a wide variety of plant species

        • Seed banks → secure sites that store and catalog seeds in order to preserve plant genetic diversity

    • A: Case study of the captive breeding and reintroduction of an endangered animal species

      • Case Study 1: Indian rhinoceros

        • An endangered species with only ca. 3500 rhinoceros living in the wild

          • This number was as low as 2000 in the early 1990s → increased due to successful captive breeding programs

        • In addition to habitat loss, the Indian rhinoceros is threatened by poachers

          • Their horn is considered potent in some cultures

        • It is listed as vulnerable on the IUCN red list + the majority of the species is situated in Indian protected areas (reserves)

      • Case Study 2: Mountain Chicken Frog

        • A species of frog native to the Carribean islands of Dominica and Montserrat

        • The population of this frog has declined i81% in the last 10 years

          • Due to the fungal disease chytridiomycosis

          • Also threatened by human consumption → local delicacy said to taste like chicken

        • Considered critically endangered with fewer than 8000 individuals existing in the wild

        • The frog has been artificially bred in laboratories in England prior to being reintroduced into the wild

  • EC: Endangered species

    • Endangered species = categorized by the International Union for Conservation of Nature (IUCN)  as likely to become extinct

      • Conservation status = likelihood of becoming extinct

    • Currently 3000 endangered species according to IUCN

    • Factors for determining a species’ conservation status

      • Rate of decline

      • Population size

      • Area of geographic distribution

      • Degree of population fragmentation

    • IUCN has a red list which classifies species into nine groups

      • Extinct (EX)

        • No surviving individuals

        • E.g. tasmanian tiger

      • Extinct in Wild (EW)

        • Captive individuals survive, but there is no wild population

        • E.g. barbary lion

      • Critically endangered (CR)

        • Faces an extremely high risk of extinction in the near future

        • E.g. red wolf

      • Endangered (EN)

        • Faces a high risk of extinction in the near future

        • E.g. snow leopard

      • Vulnerable (VU)

        • Faces a high risk of endangerment in the medium term

        • E.g. Indian rhinoceros

      • Near Threatened (NT)

        • May be considered threatened in the near future

        • E.g. tiger shark

      • Least Concern (LC)

        • No immediate threat to species survival

        • E.g. giraffe

      • All other organisms may be listed as Data Deficient (DD) or Not Evaluated (NE)

  • EC: Extinction

    • Extinction = the complete cessation of a species or higher taxon level, reducing biodiversity

    • Can occur gradually as one population of organisms evolves into something else

      • Phyletic extinction

    • Can be sudden without any identifiable descendents → cease to exits

      • Abrupt extinction

    • Can be very difficult to determine the moment of extinction → most categorizations are usually done retrospectively

      • Occasionally species thought to be extinct can be rediscovered after a period of time

        • E.g. lazarus taxa

    • 99% of all species that ever lived on Earth are considered to be extinct

      • Estimated to amount over 5 billion species

      • Natural catastrophes may cause mass extinction events

        • Large scale loss of species in relatively short period

    • Case Study: Tasmanian Tiger

      • Thylacinus cynocephalus

      • Became extinct after the arrival of European settlers to Australia

      • Tasmanian tigers would feed on introduced sheep → therefore hunted and poached

      • The loss of habitat due to human development + lack of successful breeding programs = population numbers dwindling

        • Aboriginal rock paintings suggest the Tasmanian tiger once lived on the mainland, but died out from predation and/or competition

      • Last Tasmanian tiger died in captivity in 1936

        • Declared extinct by international standards in 1986